Noyes, John Stuart, 2023, ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)

Noyes, John Stuart, 2023, ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera), Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) 2 (11), pp. 1-921

publication ID	https://doi.org/10.5281/zenodo.8074943
link to original document	https://doi.org/10.5281/zenodo.8074943
document provided by	Felipe

Treatments (399)

Hymenoptera			1
Encyrtinae			6
Encyrtidae			17-28
Encyrtidae			17
Arrhenophagus			28-29
Arrhenophagus	key		29
Arrhenophagus chionaspidis			29-30
Arrhenophagus albitibiae			30-31
Adelencyrtus			32-33
Adelencyrtus	key		33
Adelencyrtus chionaspidis			33-36
Adelencyrtus plocus		sp. nov.	36-38
Adelencyrtus odonaspidis			38-40
Adelencyrtus bifasciatus			40-41
Plagiomerus			41-42
Plagiomerus	key		42-43
Plagiomerus bangaloriensis			43-45
Plagiomerus hospes			45-49
Plagiomerus calabes		sp. nov.	49-50
Plagiomerus peruviensis			51-53
Homalopoda			53-54
Homalopoda	key		54-55
Homalopoda onothrix		sp. nov.	55-57
Homalopoda aesira		sp. nov.	57-60
Homalopoda cristata			60-61
Homalopoda rehdera		sp. nov.	61-63
Homalopoda aibeus		sp. nov.	63-65
Homalopoda beatriz		sp. nov.	65-67
Homalopoda carpha		sp. nov.	67-69
Homalopoda baunos		sp. nov.	69-70
Homalopoda opsis		sp. nov.	70-72
Homalopoda donatos		sp. nov.	72-74
Pseudhomalopoda			74-75
Pseudhomalopoda	key		75
Pseudhomalopoda tapura		sp. nov.	75-77
Pseudhomalopoda festa		sp. nov.	77-78
Pseudhomalopoda prima			79
Caenohomalopoda			80-82
Coccidencyrtus			82-84
Caenohomalopoda guamensis			82
Coccidencyrtus	key		84-85
Coccidencyrtus pomadus		sp. nov.	86-88
Coccidencyrtus viviana		sp. nov.	88-89
Coccidencyrtus aries		sp. nov.	89-91
Coccidencyrtus gowanus		sp. nov.	91-92
Coccidencyrtus reshan		sp. nov.	93-94
Coccidencyrtus juno		sp. nov.	94-96
Coccidencyrtus chenoa		sp. nov.	96-97
Coccidencyrtus procris		sp. nov.	97-99
Coccidencyrtus umbro		sp. nov.	99-101
Coccidencyrtus cyathus		sp. nov.	101-103
Coccidencyrtus xero		sp. nov.	103-105
Coccidencyrtus xylomus		sp. nov.	105-107
Coccidencyrtus jacintha		sp. nov.	107-109
Coccidencyrtus sivan		sp. nov.	109-110
Coccidencyrtus ochraceipes			110-111
Coccidencyrtus quebrus		sp. nov.	112-113
Coccidencyrtus fescus		sp. nov.	113-115
Coccidencyrtus tristeryx		sp. nov.	115-116
Coccidencyrtus matias		sp. nov.	116-118
Coccidencyrtus harpales		sp. nov.	118-120
Coccidencyrtus solivus		sp. nov.	120-121
Coccidencyrtus thuryus		sp. nov.	121-123
Quetsira		gen. nov.	123-124
Quetsira tinthrax		sp. nov.	124-125
Neococcidencyrtus			126-127
Neococcidencyrtus	key		127-128
Neococcidencyrtus mansua		sp. nov.	128-129
Neococcidencyrtus grias		sp. nov.	130-131
Neococcidencyrtus drysus			131-132
Neococcidencyrtus cyclope		sp. nov.	132-133
Neococcidencyrtus hilaria		sp. nov.	134-135
Neococcidencyrtus orion		sp. nov.	135-137
Neococcidencyrtus crouzelae			137-138
Neococcidencyrtus dracaenus		sp. nov.	138-140
Neococcidencyrtus plinthus		sp. nov.	140-141
Comperiella			142-143
Comperiella lemniscata			143-144
Orixia		gen. nov.	144-145
Orixia	key		146
Orixia demos		sp. nov.	146-147
Orixia philotes		sp. nov.	147-148
Orixia sirus		sp. nov.	148-150
Cerapterocerus			150-151
Cerapterocerus phragmitis			151-152
Helegonatopus			152-154
Helegonatopus pseudophanes			154-156
Mahencyrtus			157-158
Mahencyrtus nitidus			158-159
Tico	key		160
Tico		gen. nov.	160
Tico sinabelus		sp. nov.	161-162
Tico ryaspis		sp. nov.	162-163
Georynus		gen. nov.	163-165
Georynus	key		165
Georynus aitne		sp. nov.	165-167
Georynus dionysus		sp. nov.	167-168
Prionias		gen. nov.	168-169
Prionias nodutus		sp. nov.	169-171
Strigenia		gen. nov.	171-172
Strigenia laras		sp. nov.	172-173
Stadima		gen. nov.	174
Stadima moros		sp. nov.	175-176
Oodia		gen. nov.	176-177
Oodia japetus		sp. nov.	177-178
Ludesia		gen. nov.	178-179
Ludesia	key		180
Ludesia hemera		sp. nov.	180-182
Ludesia amalthea		sp. nov.	183-185
Angebra		gen. nov.	185
Angebra	key		186
Angebra praxadike		sp. nov.	186-187
Angebra harpalyke		sp. nov.	187-189
Anisophleps			189-190
Anisophleps aoedis		sp. nov.	190-192
Anisophleps	key		190
Anisophleps carpo		sp. nov.	192-194
Lakshaphagus			194-195
Lakshaphagus arnulfoi		sp. nov.	195-197
Cheiloneurus			197-200
Cheiloneurus	key		200-204
Cheiloneurus magnolia		sp. nov.	213-215
Cheiloneurus vitara		sp. nov.	215-216
Cheiloneurus argentipes		comb. nov.	216-218
Cheiloneurus sophocles		sp. nov.	219-220
Cheiloneurus carita		sp. nov.	220-222
Cheiloneurus epytus		comb. nov.	222-223
Cheiloneurus danessa		sp. nov.	223-225
Cheiloneurus idris		sp. nov.	225-227
Cheiloneurus socrates		sp. nov.	227-228
Cheiloneurus roellus		sp. nov.	229-230
Cheiloneurus dalycera		sp. nov.	230-232
Cheiloneurus ection		sp. nov.	232-233
Cheiloneurus parvus			234
Cheiloneurus semis		sp. nov.	235-237
Cheiloneurus catreta		sp. nov.	237-239
Cheiloneurus herndonia		sp. nov.	239-240
Cheiloneurus exeres		sp. nov.	241-243
Cheiloneurus fidenas		sp. nov.	243-245
Cheiloneurus miklo		sp. nov.	245-247
Cheiloneurus lamus		sp. nov.	247-249
Cheiloneurus varablus		sp. nov.	249-251
Cheiloneurus myndus		sp. nov.	251-253
Cheiloneurus rosea		sp. nov.	253-255
Cheiloneurus salome		sp. nov.	255-257
Cheiloneurus loresa		sp. nov.	257-259
Cheiloneurus onestas		sp. nov.	259-261
Cheiloneurus mirabel		sp. nov.	262-263
Cheiloneurus mantua		sp. nov.	264-265
Cheiloneurus hamadryas		sp. nov.	265-267
Cheiloneurus mardor		sp. nov.	268-269
Cheiloneurus extillus		sp. nov.	270-271
Cheiloneurus racidia		sp. nov.	272-274
Cheiloneurus miranda		sp. nov.	274-276
Cheiloneurus scolopax		sp. nov.	276-278
Cheiloneurus genys		sp. nov.	278-280
Cheiloneurus fabius		sp. nov.	281-283
Cheiloneurus nintra		sp. nov.	283-285
Cheiloneurus santaros		sp. nov.	286-288
Cheiloneurus simus		sp. nov.	288-290
Cheiloneurus marcius		sp. nov.	290-292
Cheiloneurus castalia		sp. nov.	293-295
Cheiloneurus dipsas		sp. nov.	295-297
Cheiloneurus medusa		sp. nov.	297-299
Cheiloneurus amadeus		sp. nov.	300-302
Cheiloneurus grimus		sp. nov.	302-304
Cheiloneurus herxius		sp. nov.	304-307
Cheiloneurus seius		sp. nov.	307-309
Cheiloneurus moloch		sp. nov.	309-311
Cheiloneurus poeana		sp. nov.	311-314
Cheiloneurus oshtan		sp. nov.	314-316
Cheiloneurus munda		sp. nov.	317-319
Cheiloneurus varus		sp. nov.	319-321
Cheiloneurus iphigenes		sp. nov.	322-324
Cheiloneurus solon		sp. nov.	324-326
Cheiloneurus pometia		sp. nov.	327-329
Cheiloneurus genua		sp. nov.	329-331
Cheiloneurus goldus		sp. nov.	331-333
Cheiloneurus aliphera		sp. nov.	334-336
Cheiloneurus mago		sp. nov.	336-338
Cheiloneurus mckenziei		sp. nov.	339-342
Cheiloneurus origo		sp. nov.	342-344
Cheiloneurus argos		sp. nov.	344-346
Cheiloneurus feritas		sp. nov.	347-349
Cheiloneurus paean		sp. nov.	349-351
Cheiloneurus bion		sp. nov.	352-353
Cheiloneurus zephyrus		sp. nov.	354-356
Cheiloneurus pherus		sp. nov.	356-358
Cheiloneurus lydia		sp. nov.	358-361
Cheiloneurus parnisa		sp. nov.	361-363
Cheiloneurus horocles		sp. nov.	363-366
Cheiloneurus longisetaceus		stat. nov.	366-367
Cheiloneurus numida		sp. nov.	368-370
Cheiloneurus aunes		sp. nov.	370-372
Cheiloneurus elegans			372-374
Cheiloneurus tainus			374-376
Cheiloneurus leda		sp. nov.	377-379
Cheiloneurus luceres		sp. nov.	379-381
Cheiloneurus perses		sp. nov.	381-383
Cheiloneurus albinotatus			383-384
Cheiloneurus dymorus		sp. nov.	384-386
Cheiloneurus glycon		sp. nov.	386-388
Cheiloneurus meryae		sp. nov.	389-391
Cheiloneurus zeno		sp. nov.	391-393
Cheiloneurus cupreicollis		stat. nov.	393-395
Cheiloneurus repola		sp. nov.	395-397
Cheiloneurus compressicornis			397-399
Cheiloneurus leone		sp. nov.	399-401
Cheiloneurus betes		sp. nov.	401-403
Cheiloneurus diaces		sp. nov.	403-406
Cheiloneurus ariston		sp. nov.	406-408
Cheiloneurus jeroba		sp. nov.	408-410
Cheiloneurus letos		sp. nov.	410-413
Cheiloneurus cero		sp. nov.	413-415
Cheiloneurus finator		sp. nov.	415-417
Cheiloneurus idifex		sp. nov.	418-420
Cheiloneurus orbona		sp. nov.	420-422
Cheiloneurus fautrix		sp. nov.	422-425
Cheiloneurus aesernia		sp. nov.	425-427
Cheiloneurus gyges		sp. nov.	427-429
Cheiloneurus lexovus		sp. nov.	429-431
Cheiloneurus numitor		sp. nov.	432-434
Cheiloneurus mora		sp. nov.	434-436
Cheiloneurus ceiba		sp. nov.	436-438
Cheiloneurus nasica		sp. nov.	439-441
Cheiloneurus alopes		sp. nov.	441-444
Cheiloneurus janzeni		sp. nov.	444-446
Cheiloneurus dirce		sp. nov.	446-448
Cheiloneurus amethes		sp. nov.	448-450
Cheiloneurus metheus		sp. nov.	451-453
Cheiloneurus omorus		sp. nov.	453-455
Cheiloneurus nebucha		sp. nov.	455-457
Prochiloneurus			458-459
Prochiloneurus	key		459
Prochiloneurus surwaka		sp. nov.	461-463
Prochiloneurus alasirus		sp. nov.	463-465
Prochiloneurus olax		sp. nov.	465-466
Prochiloneurus armidus		sp. nov.	467-468
Prochiloneurus akea		sp. nov.	469-470
Prochiloneurus abelio		sp. nov.	470-473
Prochiloneurus helleria		sp. nov.	473-475
Prochiloneurus lenus		sp. nov.	475-477
Prochiloneurus aethus		sp. nov.	477-480
Prochiloneurus dactylopii			481-482
Prochiloneurus thonus		sp. nov.	482-485
Prochiloneurus lucetia		sp. nov.	485-487
Prochiloneurus zunigai		sp. nov.	488-490
Diversinervus			490-492
Diversinervus elegans			492-494
Iceromyia			495-496
Iceromyia flavifrons			496-497
Homosemion			497-498
Homosemion bennetti			498-499
Eusemion			499-500
Eusemion	key		500
Eusemion monos		sp. nov.	500-502
Eusemion themisto		sp. nov.	502-503
Anicetus			504-505
Anicetus	key		505-506
Anicetus annulatus			506-508
Anicetus arche		sp. nov.	508-510
Anicetus oroso		sp. nov.	510-511
Anicetus lysithea		sp. nov.	511-515
Anicetus cheras		sp. nov.	515-516
Anicetus apatos		sp. nov.	517-518
Ammonoencyrtus			518-519
Ammonoencyrtus	key		520
Ammonoencyrtus hyperion		sp. nov.	520-522
Ammonoencyrtus prosymnos		sp. nov.	522-524
Ammonoencyrtus carolinensis			524-525
Ammonoencyrtus keira		sp. nov.	526-527
Ammonoencyrtus galas		sp. nov.	527-529
Ammonoencyrtus xanthos		sp. nov.	529-531
Ammonoencyrtus autone		sp. nov.	531-532
Polaszekius		gen. nov.	532-533
Polaszekius nemesis		sp. nov.	533-535
Protyndarichoides			535-536
Protyndarichoides	key		536-537
Protyndarichoides hermippe		sp. nov.	537-538
Protyndarichoides isonus		sp. nov.	538-540
Protyndarichoides nigriceps			540-541
Protyndarichoides elathes		sp. nov.	541-543
Protyndarichoides euanthos		sp. nov.	543-544
Protyndarichoides damocles		sp. nov.	545-546
Protyndarichoides hephaestus		sp. nov.	546-548
Gahaniella			548-549
Gahaniella	key		549
Gahaniella chaldenes		sp. nov.	549-551
Gahaniella californica			551-552
Gahaniella saissetiae			552-553
Prionomitus			553-554
Prionomitus mitratus			554-555
Sectiliclava			555-556
Sectiliclava	key		556
Sectiliclava isis			557
Sectiliclava helikes		sp. nov.	558-559
Sectiliclava megleos		sp. nov.	559-561
Sectiliclava fusui		sp. nov.	561-563
Sectiliclava arion		sp. nov.	563-564
Sectiliclava pulchriceps			564-565
Sectiliclava rojasorum		sp. nov.	565-567
Trechnites			567-568
Trechnites	key		568-569
Trechnites merops			569-570
Trechnites thelxines		sp. nov.	570-572
Trechnites thyones		sp. nov.	572-573
Trechnites kale		sp. nov.	573-575
Trechnites spondes		sp. nov.	575-576
Psyllaephagus			577-578
Psyllaephagus	key		579-582
Psyllaephagus domitius			582-583
Psyllaephagus ufens			583-584
Psyllaephagus heles		sp. nov.	584-585
Psyllaephagus alisanos		sp. nov.	586-588
Psyllaephagus othrys			586
Psyllaephagus trioziphagus			588-589
Psyllaephagus creusa			589-590
Psyllaephagus kallichoa		sp. nov.	590-592
Psyllaephagus epulo			592-593
Psyllaephagus pilosus			593-594
Psyllaephagus yaseeni			594-596
Psyllaephagus xepo		sp. nov.	596-598
Psyllaephagus thonis			598-599
Psyllaephagus alexion			599
Psyllaephagus rotundiformis			600-601
Psyllaephagus korus		sp. nov.	601-602
Psyllaephagus moros		sp. nov.	603-605
Psyllaephagus tyche			603
Psyllaephagus phelys		sp. nov.	605-607
Psyllaephagus gyces			607-608
Cercobelus			608-609
Cercobelus	key		609-610
Cercobelus epimetheus		sp. nov.	611-613
Cercobelus adrastea		sp. nov.	613-614
Cercobelus isara			614-615
Cercobelus sithon			615-616
Cercobelus pleiades		sp. nov.	616-617
Cercobelus tartarus		sp. nov.	617-619
Cercobelus ulixes			619-620
Cercobelus phanes		sp. nov.	620-621
Cercobelus godoyae			621-622
Cercobelus daphne			622
Arianus		gen. nov.	622-623
Arianus porusos		sp. nov.	624-625
Caldencyrtus			625-626
Caldencyrtus	key		627
Caldencyrtus balor		sp. nov.	628-629
Caldencyrtus meresmo		sp. nov.	629-631
Caldencyrtus olwen		sp. nov.	631-632
Caldencyrtus acamas			632-633
Caldencyrtus antander			633-634
Caldencyrtus mitchelli			634-635
Dalek		gen. nov.	635-637
Dalek	key		637-638
Dalek thanatos		sp. nov.	639-640
Dalek carme		sp. nov.	640-642
Dalek ananke		sp. nov.	642-644
Dalek orthia		sp. nov.	644-646
Dalek pleione		sp. nov.	646-648
Dalek jigas		sp. nov.	648-650
Dalek minerva		sp. nov.	650-652
Dalek arax		sp. nov.	652-654
Dalek saurax		sp. nov.	654-656
Dalek cyclos		sp. nov.	656-658
Dalek gubrana		sp. nov.	658-660
Dalek pomona		sp. nov.	660-662
Dalek elana		sp. nov.	662-664
Dalek nationi		sp. nov.	664-666
Mendisa		gen. nov.	666-667
Mendisa gaia		sp. nov.	667-668
Harlarus		gen. nov.	669
Harlarus titana		sp. nov.	670-671
Raquanus		gen. nov.	671-672
Raquanus hades		sp. nov.	672-674
Esphora		gen. nov.	674-675
Esphora	key		675
Esphora cyllene		sp. nov.	675-677
Esphora rhea		sp. nov.	677-679
Sosara		gen. nov.	679-680
Sosara geras		sp. nov.	680-681
Hebynthus		gen. nov.	682-683
Hebynthus	key		683-685
Hebynthus kapo		sp. nov.	685-686
Hebynthus maros		sp. nov.	687-688
Hebynthus phyrra		sp. nov.	689-690
Hebynthus cleon		sp. nov.	690-692
Hebynthus solex		sp. nov.	692-694
Hebynthus himalia		sp. nov.	694-696
Hebynthus cosmus		sp. nov.	696-698
Hebynthus menai		sp. nov.	698-700
Hebynthus cira		sp. nov.	700-701
Hebynthus erythrax		sp. nov.	702-704
Hebynthus callirhoe		sp. nov.	704-706
Hebynthus tamones		sp. nov.	706-708
Alkonia		gen. nov.	708
Alkonia tiquanus		sp. nov.	709-710
Elaxis		gen. nov.	710-711
Elaxis auster		sp. nov.	711-712
Index			713-733

Copyright notice

No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.

Treatments (399)

Copyright notice

Specimens

Downloads & Links