Oodia japetus, Noyes, 2023

Noyes, John Stuart, 2023, ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera), Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) 2 (11), pp. 1-921 : 177-178

Oodia japetus sp.nov.

( Figs 447-453)

Female (holotype): length, including ovipositor, 1.53mm; excluding ovipositor, 1.48mm ( CPD) .

Head generally black, frontovertex with a weak metallic, dark blue and green sheen mixed with some purple; area above top of scrobes with a dull purple sheen, area between eye and scrobe coppery purple and brassy, area immediately below eye with a relatively strong brassy and coppery lustre; scrobes with a weak metallic dark blue sheen above top of interantennal prominence, this continued weakly towards eye; interantennal prominence dorsally with a weak purple sheen, slightly greenish and brassy between and below toruli; mouth margin and lower gena coppery purple; temple and posterior part of gena relatively dull mixed coppery purple, brassy and metallic green; radicle pale orange, marked brown; scape pale orange, dorsal margin pale brown; pedicel brown, ventrally pale orange; flagellum mostly dark brown, F1-F4 ventrally pale orange, F5 ventrally orange-brown; mandible dark amber; maxillary palpi pale orange; frontovertex, gena and interantennal prominence with relatively sparse, scattered, inconspicuous brown setae; thorax generally dark brown to black; pronotum with a weak brassy and coppery sheen; mesoscutum a moderate, metallic, dark blue sheen mixed with some purple, posterior margin narrowly coppery purple; tegula orange-brown with a slight brassy sheen; axilla and anterior 0.7X of scutellum with a weak coppery purple and brassy sheen, sides and apical 0.3X of scutellum with a distinct metallic brassy and dark green sheen; metanotum and mesopleuron dark purple-brown with a slight brassy sheen; setae on dorsum of thorax dark brown and quite conspicuous; legs, including coxae, pale orange; wings mainly hyaline, but fore wing ( Fig. 451) with apex infuscate in the form a wedge-shaped mark; propodeum slightly shining, dark purple-brown, pale orange anteriorly near spiracle, metallic green and copper on side and submedially, about 12 setae outside spiracle, these not extending down side towards hind coxa; gaster largely dark brown, but Gt1 dorsally pale orange, anterolaterally mixed brassy, coppery, green and blue, sternites pale orange, hypopygium translucent, dark brown areas with a brassy and coppery purple sheen with some purple, green and blue reflections; gonostylus dark brown.

Head ( Fig. 448) about 3.8X as wide as frontovertex, in profile, about 1.8X as high as deep; inner eye margins slightly diverging in front of anterior ocellus; ocelli forming an angle of about 55°; frontovertex with evenly spaced piliferous punctures along inner eye margins and 7 or 8 below anterior ocellus, punctures moderately deep and each about 1.5-2X as wide as eye facet; ocellar area with moderately deep, fairly regular, polygonally reticulate sculpture of mesh size subequal to eye face, sculpture in front of anterior ocellus more irregular and of slightly larger mesh; temple and gena virtually smooth, but with shallow, irregular, longitudinally elongate, polygonally reticulate sculpture; scrobes shallow, well defined, meeting, with shallow, polygonally reticulate sculpture; interantennal prominence rounded dorsally, with shallow, regular, polygonally reticulate sculpture; antenna as in Fig. 447; scape hardly broadened, widest apically, about 4.3X as long as broad; F1 longest, F6 broadest, segments gradually increasing in width and decreasing in length distad, all at least a little longer than broad, linear sensilla present on all funicle segments; clava hardly broadened, about as wide as F6, a little shorter than F4-F6 combined; clypeal margin very slightly and evenly concave. Relative measurements: HW 79, HH 75, FV 21, POL 8, OOL 1, OCL 4.5, AOL 9, EL 52, EW 39, MS 27.5, SL 39, SW 9.

Thorax ( Fig. 452) with slightly transversely elongate, polygonally reticulate sculpture on pronotum that is generally of similar mesh size, but conspicuously shallower than that on frontovertex; mesoscutum with polygonally reticulate to imbricate-reticulate sculpture that is of slightly larger mesh size than that on frontovertex and clearly shallower; axilla with similar sculpture to mesoscutum, but slightly deeper; scutellum with conspicuously deeper and coarser sculpture than that on mesoscutum, but of similar mesh size, apical 0.3X and side with progressively shallower sculpture; visible part of mesoscutum about 1.9X as broad as long; scutellum about 1.1X as long as broad; setae on mesoscutum moderately dense, separated by a little less than their own lengths and of more or less uniform length, those on scutellum of similar density, but clearly longer especially towards apex; fore wing with venation and setation as in Figs 450, 451; propodeum ( Fig. 452) medially with some irregular, very shallow sculpture medially in posterior half; setae outside spiracle extending towards posterior margin, but not down side. Relative measurements: FWL 114.5, FWW 44.5; HWL 82, HWW 20.

Gaster very slightly shorter than thorax with hypopygium ( Fig. 449) reaching about 0.6X towards apex; syntergum about 0.6X as long as mid tibia; ovipositor ( Fig. 453) slightly exserted, the exserted part about 0.2X as long as mid tibial spur or a little less than 0.1X as long as mid tibia. Relative measurements: OL 50.5, GL 13 [MT 47].

Variation. Only holotype examined.

Male. Unknown.

DISTRIBUTION. Costa Rica.

HOSTS. Unknown.

MATERIAL EXAMINED.

Type material. Holotype E: COSTA RICA, Puntarenas, Est. Biol. Monteverde, 10°20’N 84°49’W, 1540m, 18-24.ii.2004 (C. Hansson) . Holotype in NHMUK .