Harlarus titana, Noyes, 2023

Noyes, John Stuart, 2023, ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera), Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) 2 (11), pp. 1-921 : 670-671

Harlarus titana sp.nov.

(Figs 1840-1846; Hab. E 274)

Female (holotype): length, 1.28mm ( CPD) .

Head with occiput shining, metallic green, frontovertex mainly metallic blue-green with a slight purple sheen between posterior ocelli and on occipital margin, face dark coppery purple immediately above scrobes, in scrobal area, on interantennal prominence and on temple and gena, area immediately below eye mixed blue, brassy and purple; mouth margin rainbow-like, narrowly margined purple, coppery, blue, green, brassy and coppery purple; frontovertex, gena and interantennal prominence with very inconspicuous translucent, white setae; antenna (Fig. 1841) with radicle dark brown; scape dark brown with a weak brassy and purple sheen, pale orange at extreme apex; pedicel dark brown, pale orange at apex and ventrally; funicle pale orange, proximal segments a little dusky; clava pale brown; maxillary palpus off-white; pronotum black, generally with a weak green and coppery sheen; propleuron and prosternum black; mesoscutum moderately bright, metallic, blue-green; tegula dark brown with a weak coppery purple sheen; axilla dark purple-brown with a weak brassy sheen; scutellum less shiny than mesoscutum but mixed dark metallic green and blue-green with some coppery purple reflections, side and apex shining dark green and blue-green; mesoscutum, clothed with fairly conspicuous, off-white, translucent setae, scutellum with conspicuous dark brown setae; metanotum dark purple-brown with a weak brassy sheen; mesopleuron dark brown, mainly with a weak brassy and coppery sheen mixed with some green and blue-green; mesosternum dark brown; all coxae dark brown; fore femur dark brown with apices very pale yellow, tibia very pale yellow with a broad, subbasal brown ring, tarsus very pale, dusky yellow, darker towards apex; mid femur very pale yellow at apices otherwise dark brown, tibia very pale yellow with a broad, dark brown, subbasal band, spur and tarsus very pale yellow, distal segments of tarsus slightly darker; hind femur dark brown with extreme apices very pale yellow, tibia dark brown with extreme base very pale yellow and apex pale orange, tarsus dark brown; wings hyaline, but fore wing with a small infuscate area adjacent to stgmal and marginal veins (Fig. 1844); propodeum dark brown with a weak brassy sheen, side with a strong, metallic blue-green sheen, about 15 conspicuous, translucent, silvery setae behind and outside spiracle extending someway down side of propodeum and towards posterior margin; gaster dark brown, with a distinct coppery purple sheen, dorsal part of Gt1 with a strong metallic blue-green sheen, side of gaster with a mixed metallic green, brassy and coppery sheen; gonostylus dark brown.

Head (Fig. 1843) about 2.7X as wide as frontovertex; ocelli forming an angle of about 100°; frontovertex with fairly regular, moderately deep, polygonally reticulate sculpture of mesh size generally subequal to eye facet, piliferous punctures shallow, 8 in front of anterior ocellus, a line of sparse setae along inner eye margin continued along lower eye margin; interantennal prominence with two submedian lines of longer, more conspicuous setae; temple and gena about as shiny as frontovertex, temple and gena with fairly shallow, longitudinally elongate, polygonally reticulate sculpture; eye virtually reaching occipital margin, separated by about 0.5X diameter of facet, with dense, inconspicuous pale setae that are each much shorter than diameter of a facet, separated from scrobal margin by about diameter of ocellus, area between eye and scrobe with transversely elongate, imbricate-reticulate sculpture; scrobes dorsally separated by interantennal prominence, very weakly margined dorsally and laterally; interantennal prominence shallow, dorsally acute and meeting frontovertex, with very shallow, polygonally reticulate sculpture; antenna as in Fig. 1841; scape strongly broadened, only about 1.7X as long as broad; clava about 1.1X as broad as F6, and about as long as F4-F6 combined, sutures very slightly oblique, sensory area, extending about 0.2X along ventral surface of clava; malar sulcus present, indistinct. Relative measurements: HW 90, HH 77, FV 33, POL 19, OOL 2, OCL 3, AOL 11, EL 51, EW 40.5, MS 30, SL 43, SW 26.

Thorax (Fig. 1846) in dorsal view with pronotum short, largely hidden by head; pronotum with similar polygonally reticulate sculpture to that on frontovertex; mesoscutum with slightly deeper, more regular, polygonally reticulate sculpture of smaller mesh; axilla with polygonally reticulate sculpture that is conspicuously shallower than that on mesoscutum; dorsal part of scutellum with longitudinally elongate polygonally reticulate to striate reticulate sculpture that is conspicuously finer and deeper than that on mesoscutum, apex and side almost smooth, but with very shallow, polygonally reticulate sculpture and relatively shiny; visible part of mesoscutum about 1.7X as broad as long; scutellum dorsally only slightly convex, about as long as broad, apex not thin and flange-like; fore wing with venation and distribution of setae as in Figs 1844, 1845; costal cell with a single line of setae dorsally in apical two-fifths or so; mid tibial spur slightly shorter than basitarsus; propodeum (Fig. 1846) medially about 0.16X as long as scutellum and more or less smooth, side with some shallow, polygonally reticulate sculpture; side of propodeum rounded posteriorly above hind coxa and without a tooth. Relative measurements: FWL 99, FWW 42; HWL 69.5, HWW 19.5.

Gaster with hypopygium reaching about 0.8X to apex; syntergum about 0.6X as long as mid tibia, with apex weakly angular, more or less rounded; ovipositor not exserted.

Paratype. Hypopygium Fig. 1840; ovipositor Fig. 1842. Relative measurements: OL 62, GL 12 [MT 77, MTS 22].

Variation. Females vary in length from 1.24-1.38mm, the dorsal part of the interantennal prominence may not quite reach the frontovertex so that the scrobes shallowly meet dorsally, otherwise very little in material available.

Male. Unknown.

DISTRIBUTION. Costa Rica.

HOSTS. Unknown.

MATERIAL EXAMINED.

Type material. Holotype E: COSTA RICA, San José , Zurqui de Moravia, 1600m, iii.1990 (P. Hanson) . Paratypes: COSTA RICA, 2E, San José, same data as holotype but iv.1991 and iv.1992 . Holotype in MZUCR, paratypes in NHMUK .