Arianus porusos, Noyes, 2023

Noyes, John Stuart, 2023, ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera), Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) 2 (11), pp. 1-921 : 624-625

Arianus porusos sp.nov.

(Figs 1694-1701; Hab. E 252)

Female (holotype): length, including ovipositor, 2.72mm; excluding ovipositor, 2.64mm (CPD).

Body generally dark brown with a weak to moderate metallic sheen; frontovertex generally with a moderately metallic dark blue-green sheen, but distinctly coppery violet when viewed from some angles, bottoms of piliferous punctures mostly coppery violet; temple and gena somewhat coppery, anterior part of gena mixed metallic dark blue-green; scrobal area mainly metallic dark blue-green but mixed coppery violet as frontovertex, interantennal prominence quite strongly violet in part, mouth margin dull coppery; antenna (Fig. 1697) with radicle pale orange with some dark brown dorsally; scape pale orange; pedicel brown, paler distally and ventrally; flagellum orange-brown; palpi pale orange, slightly darker proximally; mandible dark brown, teeth slightly paler; pronotum with a slight coppery sheen; mesoscutum predominantly with a moderate coppery sheen mixed with some very dull metallic dark green, viewed from some angles almost completely dull metallic dark green; axilla similar to mesoscutum but more strongly coppery; scutellum with a moderate coppery sheen, stronger towards apex; metanotum dark orange-brown with a slight weak coppery sheen; tegula slightly coppery; prepectus with a slight coppery sheen; mesopleuron anteriorly with a slight brassy lustre, posterior half more coppery; all coxae dark brown; fore femur dark brown with apical one-third pale orange, tibia and tarsus pale orange, apical tarsomere partially orange-brown; mid femur dark brown with apical half pale orange, tibia and tarsus pale orange, spur dark orange-brown; hind femur dark brown with apical half pale orange, tibia and tarsus pale orange; fore wing (Fig. 1701) generally very weakly infuscate with a small, posterior, dark brown, triangular area towards base, hind wing hyaline, venation pale orange; propodeum slightly coppery with a weak brassy sheen; gaster dorsally with a distinct coppery sheen with green and violet reflections, side and venter similar but less coppery; gonostylus pale brown, apex off-white.

Head (Fig. 1695) about 2.2X as wide as frontovertex, in profile about 2.1X as high as deep, frontovertex fairly evenly and slightly convex between occipital margin and top of scrobes, below top of scrobes hardly convex to mouth margin, tangent of frontovertex forming an angle of about 118° with that of face, interantennal prominence hardly protuberant between toruli; ocelli forming an angle of about 98°; frontovertex moderately shiny with fine, polygonally reticulate sculpture of mesh size slightly smaller than eye facet; piliferous punctures only slightly smaller than ocellus and very nearly touching, punctures extending partway between scrobe and eye; setal sockets on gena fine, separated by a little less than their own diameters; temple mostly with moderately deep longitudinally elongate polygonally reticulate sculpture, this continuing on to gena but anterior part of gena with more regular polygonally reticulate sculpture; antenna as in Fig. 1697; scape about 4.3X as long as broad; funicle with F1 about 1.5X and F6 about 1.3X as long as broad respectively, hardly becoming shorter or broader distally, linear sensilla present on all segments; clava 3-segmented, both sutures oblique, about 1.3X as wide as F6, about 2.8X as long as broad, slightly shorter than F4-F6 combined, sensory area extending slightly more than half way along ventral surface; eye reaching occipital margin, clothed in very inconspicuous, very sparse setae, each not more than about 0.5X as long as diameter of facet. Relative measurements: HW 187.5, HH 179, FVL 110, FV 86, POL 57, OOL 3, OCL 10, AOL 32.5, EL 111.5, EW 76, MS 53, SL 58, SW 13.5.

Thorax (Fig. 1696) with pronotum exposed behind head, posterior margin hardly concave, the exposed part medially about as long as AOL; pronotum with moderately deep polygonally reticulate to coriaceous sculpture that is slightly shallower than sculpture on frontovertex; mesoscutum mostly with fine polygonally reticulate sculpture that is about as deep as that on frontovertex but generally of slightly larger mesh; axilla with slightly shallower, finer, polygonally reticulate sculpture than that on mesoscutum; scutellum with similar sculpture to mesoscutum but very slightly shallower, more irregular and of smaller mesh; visible part of mesoscutum nearly 2X as broad as long; scutellum very slightly broader than long; anterior half of mesopleuron with shallow, very poorly defined, polygonally reticulate sculpture, posterior half with distinct, well-defined, polygonally reticulate sculpture; fore wing with venation and setation as in Figs 1694, 1701; propodeum (Fig. 1696), medially with some irregular polygonally reticulate sculpture, area outside spiracle with about 20-25 conspicuous setae, these extending part way down side towards hind coxa; diameter of spiracle about equal to anterior ocellus. Relative measurements: FWL 190, FWW 79.5; HWL 118, HWW 38.

Gaster with hypopygium more or less reaching apex; syntergum about 0.7X as long as mid tibia, with apex narrowly truncate; ovipositor not exserted.

Paratype. Gaster with Gs2 as in Fig. 1698; hypopygium Fig. 1699; ovipositor Fig. 1699. Relative measurements: OL 58.5, GL 6 [MT 70].

Variation. Females vary in overall length from 2.33-2.72mm.

Male. Unknown.

HOSTS. Unknown.

DISTRIBUTION. Costa Rica.

MATERIAL EXAMINED.

Type material. Holotype E: COSTA RICA, Cartago, La Cangreja , 1950m, vii.1991 (P. Hanson, C. Godoy) . Paratypes: COSTA RICA, 1E, Alajuela, Peñas Blancas , 9.vi.1987 (E. Cruz) . Holotype in MZUCR, paratype in NHMUK .