Duarte, Paulo R. M., Grossi, Paschoal C. & Dupuis, Fabien, 2022, A taxonomic revision of the Bothynus villiersi Endrödi, 1968 species group (Coleoptera: Scarabaeidae: Dynastinae)

Duarte, Paulo R. M., Grossi, Paschoal C. & Dupuis, Fabien, 2022, A taxonomic revision of the Bothynus villiersi Endrödi, 1968 species group (Coleoptera: Scarabaeidae: Dynastinae), Zootaxa 5093 (1), pp. 49-66

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