Tomopterna tuberculosa ( Boulenger, 1882 )

Willems, Frank & Channing, Alan, 2023, The status of Tomopterna pulchra Boulenger, 1896 (Anura: Pyxicephalidae), Zootaxa 5374 (3), pp. 361-389 : 376

publication ID

https://doi.org/ 10.11646/zootaxa.5374.3.3

publication LSID

lsid:zoobank.org:pub:4FEAC6E6-32A8-4F01-BC7F-C85D11431936

DOI

https://doi.org/10.5281/zenodo.10248465

persistent identifier

https://treatment.plazi.org/id/03F0FD2E-FFEE-FFFC-FF0D-FE461CF1FA20

treatment provided by

Plazi

scientific name

Tomopterna tuberculosa ( Boulenger, 1882 )
status

 

Tomopterna tuberculosa ( Boulenger, 1882) View in CoL

Western Beaded Sand Frog

Pyxicephalus rugosus Ģnther, 1865:479.

Rana tuberculosa Boulenger, 1882:30 .

Rana (Tomopterna) signata Ahl, 1927:43 View in CoL . Synonymy by Laurent (1954).

Rana (Tomopterna) cacondana Ahl, 1927:44 . Synonymy by Laurent (1954).

Pyxicephalus tuberculosus View in CoL — Parker 1936:100.

Rana (Tomopterna) tuberculosa — Monard 1937:44.

Tomopterna cacondana View in CoL — Deckert 1938:142.

Tomopterna rugosa View in CoL — Laurent 1954:72.

Tomopterna tuberculosa View in CoL — Laurent 1964:133.

Tomopterna (Tomopterna) tuberculosa View in CoL — Dubois 1987:57.

Tomopterna tuberculosa ( Deckert, 1938) View in CoL — Lobón-Rovira et al. (2022). Attributed to Deckert (1938) in error.

Tomopterna tuberculosa View in CoL was described by Ģnther (1865) as Pyxicephalus rugosus View in CoL based on two specimens collected by Dr. Welwitsch from Pungo Andongo in Angola. Pyxicephalus rugosus View in CoL was replaced by Boulenger (1882), as it was preoccupied by Rana rugosa Temminck & Schlegel, 1838 View in CoL . In the description, Ģnther emphasises the glandular folds that commence behind each superciliary, converging toward the midline before curving outwards behind the neck. His detailed illustration ( Fig. 2 View FIGURE 2 ) shows the rough dorsum and the characteristic contrasting pattern.

Laurent synonymised T. signata ( Ahl, 1927) View in CoL and T. cacondana ( Ahl, 1927) View in CoL as the specimens were collected near the type locality of T. tuberculosa View in CoL and were essentially indistinguishable from that species.

Advertisement calls from Ongongo and Cuima have a mean dominant frequency of 2639 Hz (2608–2670 Hz), with a mean note rate of 10.5 s- 1 (8.6– 12.3 s- 1) n=2, at an ambient temperature of 24°C.

Sequenced specimens attributable to T. tuberculosa do not possess a pale vertebral stripe, but in body proportions and tubercles are similar to T. pulchra View in CoL . Poynton & Broadley (1985) noted that some Angolan specimens were brown overall, rather than patterned dorsally. At Ongongo in Namibia about 80% of calling males were brown, with the remainder showing the highly contrasting pattern. One of each pattern is illustrated in Fig. 2 View FIGURE 2 .

Tomopterna tuberculosa has only been genetically confirmed from western Angola and northwest Namibia ( Fig. 7 View FIGURE 7 ).

Kingdom

Animalia

Phylum

Chordata

Class

Amphibia

Order

Anura

Family

Pyxicephalidae

Genus

Tomopterna

Loc

Tomopterna tuberculosa ( Boulenger, 1882 )

Willems, Frank & Channing, Alan 2023
2023
Loc

Tomopterna (Tomopterna) tuberculosa

Dubois, A. 1987: 57
1987
Loc

Tomopterna tuberculosa

Laurent, R. F. 1964: 133
1964
Loc

Tomopterna rugosa

Laurent, R. F. 1954: 72
1954
Loc

Tomopterna cacondana

Deckert K. 1938: 142
1938
Loc

Rana (Tomopterna) tuberculosa

Monard, A. 1937: 44
1937
Loc

Pyxicephalus tuberculosus

Parker, H. W. 1936: 100
1936
Loc

Rana (Tomopterna) signata

Ahl, E. 1927: 43
1927
Loc

Rana (Tomopterna) cacondana

Ahl, E. 1927: 44
1927
Loc

Rana tuberculosa

Boulenger, G. A. 1882: 30
1882
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