Titiotus marin, Platnick & Ubick, 2008

Platnick, Norman I. & Ubick, Darrell, 2008, A Revision of the Endemic Californian Spider Genus Titiotus Simon (Araneae, Tengellidae), American Museum Novitates 3608 (1), pp. 1-36 : 30-33

Titiotus marin , new species

Figures 77–81 ; map 3

TYPE: Male holotype from Mill Valley , Marin Co. , California (June 1951; D. Hardy), deposited in AMNH.

ETYMOLOGY: The specific name is a noun in apposition taken from the type locality.

DIAGNOSIS: Males resemble those of T. humboldt and T. costa but can be distinguished by having the third most ventral prong of the tibial apophysis notched (fig. 79); females have a relatively narrow, ovoid epigynal septum (fig. 80).

MALE: Total length 11. Coloration as in T. californicus . Leg spination: femora: I p0-3-1, r1-2-1; II r0-2-1; III r1-1-2; IV r0-1-2; tibiae: I, II d1-0-1, v4-4-4, r1-1-0; III r0-1-1; metatarsi: I, II p1-1-0, r1-1-0; III p1-2-2; IV r1-2-1. Embolus prolaterally expanded near tip, distally narrow; median apophysis with only tiny subdistal projection; most ventral prong of tibial apophysis triangular, middle prong relatively short, distally incised, dorsal prong distally narrowed (figs. 77–79).

FEMALE: Total length 11. Coloration as in T. californicus . Leg spination: femora: I–III r1-1-2; IV p1-2-1, r0-1-2; tibiae: I p1-0-0, v4-4- 4, r1-0-0; II d1-0-0, p0-1-1, v4-4-4, r1-1-0; III r0-1-1; metatarsi III p1-2-2. Epigynal septum relatively narrow, ovoid, posterolateral epigynal margins protuberant; posterior ducts oblique (figs. 80, 81).

OTHER MATERIAL EXAMINED: United States: CALIFORNIA: Marin Co. : Burdell Mountain, SE slope, Novato , 38 ° 07 9 48 0 N, 122 ° 34 9 09 0 W, Mar. 16, 1990, serpentine grassland with adjacent mixed woodland (D.

Ubick, T. Briggs, CDU), 13, Mar. 8, 1991, oak woodland, under serpentine (D. Ubick, CDU), 1♀; Fairfax , Nov. 15, 1974 ( CAS) , 13, 38 ° 00.0 9 N, 122 ° 36.0 9 W, May 6, 1999, in house in oak woodland (J. Schweikert, CAS) , 13, Mt. View Road , Apr. 28, 2004 (D. Giampietro, CAS) , 13; Marinero Estates , Tiburon Peninsula, Apr. 9, 1994, serpentine grassland (D. Ubick, CDU) , 23; Mill Valley , no date, running on floor in house (E. Ross, CAS) , 1♀, June 1951 (D. Hardy, AMNH) , 13, June 7, 1954 (H. Leech, AMNH) , 13, June 17, 1955, at light (H. Leech, AMNH) , 13, June 1956 (E. Ross, AMNH) , 13, Aug. 1956, in house (E. Ross, AMNH) , 1♀; Apr. 1957, in empty bathtub at night (E. Ross, AMNH) , 13, June 14, 1957, in house (H. Leech, AMNH) , 13, June 1, 1958 (F. Leech, CAS) , 13, Sept. 22, 1958 (H. Leech, CAS) , 13, Oct. 1, 1958, in house (E. Ross, AMNH) , 1♀, Aug. 1, 1963, in house at night (E. Ross, CAS) , 1♀, May 1965, house basement (E. Ross, CAS) , 23, 1♀, June 6, 1988, in basement of house (C. Hausner, CDU) , 1♀, Sept. 5, 1991 (L. Alderson, CDU) , 1♀, June 9, 1995 ( V. Roth , CAS), 13, June 11, 1996, in house (E. Ross, CAS) , 13, Aug. 4, 2002 (S. Hall, UCR) , 1♀, May 30, 2003 (E. May, UCR ), 1♀; Mount Tamalpais , Mar. 14, 1909 (E. VanDyke, CAS) , 13; Novato , May 6, 1973 (B. Kessel, CAS) , 13, Mar. 14, 1982, under serpentine (D. Ubick, CDU) , 13, Apr. 25, 1982, under serpentine float (D. Ubick, CDU) , 13, Jan. 1990, under serpentine (D. Ubick, CDU) , 13; Ring Mountain , Tiburon Peninsula, Apr. 15, 1994, serpentine grassland (D. Ubick, J. Boutin, CDU) , 13 (penultimate, but palp details visible through cuticle); San Anselmo , May 22, 1998, in house ( CAS) , 1♀, Aug. 29, 2000, in house (D. Burns, CAS) , 1♀; San Geronimo , Jan. 15, 1994 (matured Mar. 24, 1994), serpentine grassland (D. Ubick, J. Boutin, CDU) , 13; San Rafael , June 28, 1993, in house (N. Durighello, CDU) , 1♀, mid-May 2000, in house (D. Halliday, UCR) , 13; N San Rafael, Feb. 6, 1988 (3 matured Mar. 21, 1988, ♀ Apr. 4, 1988), serpentine grassland (D. Ubick, CDU) , 13, 1♀; San Rafael Ridge , San Anselmo, May 27, 1996 (L. Freihofer, CAS) , 1♀; Sausalito (hilly part), May 23, 2000, in house (A. Weisskopf, CAS) , 13; South Tiburon Ridge , 37 ° 53 9 05 0 N, 122 ° 27 9 06 0 W, Feb. 27, 1993, serpentine grassland (D. Ubick, J. Boutin, C. Kellner, CDU) , 2♀, Apr. 18, 1993, serpentine grassland (D. Ubick, CDU) , 23, 1♀; Terra Linda , San Rafael, 38 ° 01.2 9 N, 122 ° 34.5 9 W, all border of suburb and open space, grassy hills, oaks, laurels, shrubs, elev. 100 m (J. Schweikert, CAS) : June 4, 2003, 13, June 22, 2003, 13, July 15, 2003, 13, July 31, 2003, 1♀, Aug. 23, 2003, 1♀, Apr. 13, 2004, 13, Apr. 25, 2004, 13, May 2, 2004, 13, July 18, 2004, 13, May 29, 2005, 13, May 13, 2006, 13; Tiburon, Dec. 21, 1988 (L. Cheng, T. Briggs, CAS) , 1♀, mid-April 2002 (K. Passaretti, UCR) , 23; Woodacre , Apr. 21, 1981, in house (P. Signet, UCB) , 13. Mendocino Co.: Ukiah , June 5, 2000, in house (E. Nicolait, CAS) , 13. Sonoma Co.: Petaluma , May 16, 2003, in bathroom (H. Stavropoulos, UCR) , 13.

DISTRIBUTION: Coastal California, from the San Francisco Bay area north (map 3) .

Figures

Figs. 77–81. Titiotus marin, new species. 77. Left male palp, prolateral view. 78. Same, ventral view. 79. Same, retrolateral view. 80. Epigynum, ventral view. 81. Same, dorsal view.