Teremitra efatensis ( Aubry, 1999 ) Aubry, 1999

Kantor, Yuri, Lozouet, Pierre, Puillandre, Nicolas & Bouchet, Philippe, 2014, Lost and found: The Eocene family Pyramimitridae (Neogastropoda) discovered in the Recent fauna of the Indo-Pacific, Zootaxa 3754 (3), pp. 239-276 : 266

Teremitra efatensis ( Aubry, 1999) comb. nov.

Figures 2 P, 15A–I, 16A–C

Terebra efatensis Aubry, 1999: 39 .

Terebra efatensis— Terryn 2007: 32, pl. 37, fig. 4, pl. 62 (holotype illustrated). Pristiterebra efatensis — Terryn & Holford 2008: 46, pl. 18, figs. 3–4.

Type material. Holotype ( MNHN 2798) and 2 paratypes ( MNHN 2800) from the type locality. Other paratypes as listed below.

Type locality. Vanuatu, 16º40'S, 168º04'E, 450–489 m [MUSORSTOM 8, st. DW 1048].

Material examined. The type material.

Philippines. 08º56.5’N, 123º32.7’E, 679–740 m, off Siquijor I., 3 dd.

Vanuatu or SW New Caledonia (accidentally mixed stations), 2 paratypes ( MNHN 2817).

Vanuatu. MUSORSTOM 8, st. DW 1045, 16º54'S, 168º20'E, 459–488 m, 1 lv (paratype, MNHN 2799); SANTO 2006, st. AT10, 15º41.1’S, 167º00.5’E, 509-659 m, W Malo Island, 2 dd.

Fiji islands. BORDAU 1, st. DW1432, 17°20'S, 178°44'W, 477–493 m, Lau Ridge, Vanua Balavu, 2 dd ( MNHN IM–2012–812).

Description. Shell small, narrow, terebriform, solid, with high elevated spire (LWL/SL 0.39-0.47) and attenuated short canal. Shell consisting of 2.5 protoconch whorls and up to 10 convex, slightly angulated, teleoconch whorls separated by distinct, impressed suture. Protoconch ( Fig. 15 F, H) diameter 640–650 Μm, height 660–700 Μm, light brown; initial 1.5–1.75 whorls rounded and convex, smooth, bordered posteriorly by thin, distinct, opisthocyrt ribs, 16 on last protoconch whorl; transition to teleoconch sharp. Axial sculpture of teleoconch consisting of nearly straight, slightly opisthocline, ribs, 20–21 on penultimate whorl, 18–20 on last whorl. Spiral sculpture consisting of three strong cords on spire whorls; adapical-most subsutural cord single on first whorls, becoming subdivided in two closely spaced ones on later whorls; last whorl with 7–8 major cords and 8–9 smaller cords on canal. Axial ribs forming beaded intersections with cords. Aperture narrowly elongate, low (AL/SL 0.32–0.34). Inner lip covered with thin, glossy callus. Columella with two weakly pronounced, oblique folds, better seen when shell is slightly turned counter-clockwise. Siphonal canal broad, weakly delimited from aperture, recurved abaxially. Shell color offwhite, slightly glossy under thin, transparent to light olive, smooth, peeling periostracum.

Measurements (holotype, the largest specimen): SL 10.8 mm; LWL 4.3 mm, AL (with canal) 3.5 mm, SW 2.9 mm.

One paratype (MNHN 2799) contained a dried out body, which was dissected after rehydration. Operculum absent. Head small, long, with pronounced neck, very short tentacles, and small eyes. Penis extremely long, spanning about one whorl, simple, sharply narrowing at about mid-length, without distinct papilla. Columellar muscle long, spanning about two whorls. Midgut gland medium-sized, partially tubular, rather thick. Proboscis long and coiled. Radula short ( Fig. 16 A–C), about 240 Μm long (9.2% of AL), consisting of about 30 rows of teeth, 10 nascent, with strong alary processes of membrane embracing anterior part of odontophore, membrane folded along both sides of the rachidians. Rachidian teeth with broad base and large cusp, sometimes with indistinct additional serration. Lateral teeth high triangular, shallowly scoop-shaped, attached by narrow base; inner side adjoining the rachidian bearing two small, narrow but rather long, additional cusps.

Distribution. The Philippines, Vanuatu and Fiji, alive in 459–488 m, shells to 679 m.

Figures

FIGURE 2. Overview of fossil and Recent Pyramimitridae. A. Endiatoma quadricincta (Cossmann, 1883), MNHN. F. A 47757, Lower Eocene, France. B – C. Pyramimitra terebraeformis (Conrad, 1848), MNHN. F. J 08748, Middle Eocene, USA. D – E. Pyramimitra quadralirata Dockery, 1977, MNHN. F. A 47750, Middle Eocene, USA. F. Hortia priabonica n. sp., MNHN. F. A 47752, Upper Eocene, France. G. Hortia arriuensis Lozouet, 1999, MNHN. F. A 47751, Upper Oligocene, France. H. Hortia pseudotaranis n. sp., New Caledonia, SL 8.4 mm. I. Hortia macrocephala n. sp., New Caledonia, SL 10.4 mm. J. Hortia paradrillia n. sp., New Caledonia, SL 8.1 mm. K. Hortia marshalli n. sp., New Zealand, SL 10.6 mm. L. Hortia spenceri n. sp., New Zealand, SL 9.0 mm. M. Hortia aotearoa n. sp., New Zealand, SL 9.0 mm. N. Hortia solitaria n. sp., South of Madagascar, SL 6.5 mm. O. Teremitra fallax n. sp., Solomon Islands, SL 11.8 mm. P. Teremitra efatensis (Aubry, 1999), Vanuatu, SL 10.8 mm. Q. Teremitra? hortensis (Vinassa de Regny, 1898), Upper Eocene, France. R. Teremitra sp., Taiwan, SL 16 mm. S. Vaughanites? superstes n. sp., Philippines, SL 19.1 mm.

FIGURE 15. Shells of Teremitra efatensis (Aubry, 1999) (A – I) and T. fallax n. sp. (J – M). Teremitra efatensis (Aubry, 1999) (A – I). A – C. Holotype (MNHN 2798), SL 10.75 mm. D – E. Paratype (MNHN 2799), Vanuatu, MUSORSTOM 8, st. DW 1045, SL 9.4 mm (radula illustrated). F – G. Paratype (MNHN 2817), Vanuatu or SW New Caledonia, SL 10.2 mm. H – I. Fiji islands, Lau Ridge, Vanua Balavu, BORDAU 1, st. DW 1432, SL 10.0 mm. Teremitra fallax n. sp. (J – M), holotype, SL 11.8 mm. All shells at the same scale.

FIGURE 16. Radulae of Teremitra (A – F) and Vaughanites? superstes n. sp. (G – H). A – C. Teremitra efatensis, paratype, Vanuatu, MUSORSTOM 8, st. DW 1045, 16 º 54 ' S, 168 º 20 ' E, 459 – 488 m, SL 9. 4 mm. A. Central part of the membrane, dorsal view. B. Enlarged part of bending plane. C. Entire radular membrane. D – F. Teremitra fallax n. sp., holotype. D. Dorsal view of the central part of the radular membrane. E. Enlarged rachidian teeth. F. Enlarged lateral teeth. G – H. Vaughanites? superstes n. sp., holotype. G. Dorsal view of the central part of the radular membrane. H. Enlarged part of bending plane to show the lateral teeth.