Ptychadena bunoderma (Boulenger, 1907)

Conradie, Werner, Keates, Chad, Verburgt, Luke, Baptista, Ninda L. & Harvey, James, 2023, Contributions to the herpetofauna of the Angolan Okavango- Cuando-Zambezi river drainages. Part 3: Amphibians, Amphibian & Reptile Conservation (e 325) 17 (1), pp. 19-56 : 42

Ptychadena bunoderma (Boulenger, 1907)

Rough Ridged Frog ( Fig. 29; Map 26)

Material (2 specimens): PEM A12778, Quembo River source, trap 4, -13.13586° 19.04709°, 1,369 m asl; PEM A12476, Cuanavale River source lake, -13.08537° 18.89100°, 1,360 m asl. Description: Small stocky Ptychadena ; dorsum warty, with no clear ridges as in other Ptychadena species; short stubby legs; very reduced webbing, four phalanges of the 4 th toe free of webbing. Dorsum dark brown to olive, with scattered white to brown elevated tubercles; lower lip gray with white bars; back of thighs with scattered light spots. The only adult male measured 25.6 mm ( PEM A12778). Habitat and natural history notes: Specimens were found in trap arrays (see Conradie et al. 2021) set up next to flooded areas of source lakes. They were never heard calling, and either have a very cryptic call or were not vocal during the survey periods. Comments: In Angola, this species is only known from the east, based on historical records ( Marques et al. 2018). No new collections were made until Ernst et al. (2020) recorded a series of specimens from northwestern Angola. Elsewhere, it is known from northwestern Zambia ( Channing 2001; Channing and Rödel 2019). Our new material represents a new record for southeastern Angola and the Okavango River basin.