Pseudoneureclipsis complexa, Oláh & Johanson, 2010

Oláh, János & Johanson, Kjell Arne, 2010, Fifteen new Trichoptera (Insecta) species from Sumatra, Indonesia, Zootaxa 2618 (1), pp. 1-35 : 7-9

Pseudoneureclipsis complexa , new species

Figs 8–11

This species is chestnut brown and has a unique complex of dorsobasal processes in the genitalia, each with 3 lobes. In other species the dorsobasal processes form simple, spine-like structures with only slight modifications.

Male. Head pale yellow except posterior part of labrum and sutures being chestnut brown; shape almost quadrangular in dorsal view. Ocelli absent. Eye diameter about half vertex width. Maxillary palp formula II-I- III-IV-V. Antennae stout, slightly shorter than forewings; scapes short, 1/4th head length; interantennal distance wider than each scape. Labrum bipartite, posterior part sclerotized, anterior part membranous. Mandible enlarged, broad, surface granulated papillose. Laciniae elongate, setiferous, curving mesad. Anterior tentorial pits located laterally, adjacent to eyes; pregenal area narrow; clypeus and frons wide. Epistomal suture connecting anterior tentorial pits detectable between clypeus and frons, frons covered with setae. Frontal grooves almost indiscernible. Clypeolabral grooves well-developed, encircling enlarged mandibles. Coronal grooves running through middle of vertex and occiput. Occipital grooves well developed. Postoccipital groove visible between occipital setal warts. Pairs of vertexal lateroantennal and vertexal ocellar compact setose warts differentiated; vertexal medioantennal and vertexal medioocellar diffuse setose warts partly fused, forming mesal band on vertex. Pair of occipital compact setose warts dominating postdorsal region of head. Postgenal compact setose warts narrow, visible in caudal and ventral view. Pair of vertexal tubercles, sensory papillae or sensilla basiconica, present posterior of crossing coronal and occipital grooves.

Cervical sclerites with long, narrow, rod-like anterior arms articulating anteriorly with elongate occipital condyles above posterior tentorial pits; fused to posterior cervical sclerites. Posterior cervical sclerites half as long as anterior sclerites; each forming triangular plate, with 1 angle reaching and articulating with prothoracic episternum and other angle producing rod-like ventral intercervical sclerites. Ventral intercervical sclerites articulating with weakly sclerotized anteromedian band of prothoracic eusternum. Cervical sclerites without setae; single, large cervical setose wart attached to dorsum of anterior rod-like sclerites near posterior sclerites. Thoracic sclerites chestnut brown with pale setal warts. Two pairs of large, rounded, pronotal compact setose warts present; lateral pronotal warts smaller than mesal pronotal warts; mesal pronotal warts situated close to each other. Pair of mesoscutal compact setose warts rounded, distinct; pair of mesoscutellar compact setose warts longitudinally elongate, slightly triangular, narrowing posteriorly, occupying most of mesoscutellum.

Spur formula 2,4,4; foreleg posteroapical spurs each half as long as anteroapical spurs; middle leg and hind leg posteroapical spurs and posterosubapical spurs 1/3rd times longer than anteroapical and anterosubapical spurs. Leg claws small, symmetrical.

Forewings 5.0 mm long; membrane and veins light brown, covered with sparce cinereus setae. Forewings each with forks I, II, III, and IV present; discoidal, median and thyridial cells closed; crossvein m-cu closing thyridial cell reaching first point of bifurcation of M; fork I smaller than fork III. Hind wings each with forks II, III, and V present; thyridial cell closed, narrow; discoidal and median cells open.

Male genitalia. Segment IX with small tergite and large sternite; tergite IX present anterior of cerci; sternite IX tall, slightly arching with convex anterior margins and concave posterior margins in lateral view, ellipsoid, anterior and posterior margins with darkly pigmented antecostae; produced posteroventrad at articulation of gonopods. Segment X produced into pigmented dorsomesal depression; dark pigmentation accentuated by microtrichiae densely covering marginal lateral rims. Cerci semicircular in lateral view, triangular in dorsal view, their posterior margins serrated by elevated tubercles, covering bases of paraprocts. Paraprocts each hinged to mesal base of corresponding cercus, each divided into 2 spine-like processes. Dorsal paraproctal processes shorter than ventral paraproctal processes; ventral paraproctal processes each obliquely notched several times along apical 1/3rd. Coxopodites broad, rod-like in lateral and ventral views. Harpagones located dorsobasally, each forming complex structure with 3 lateral lobes, and fused anteriorly with coxopodites by median ridge, together forming apodeme or basal plate. Phallic apparatus with ventradcurving, basally broadening phallobase; continuing into heavily sclerotized, ventrally tapering process. Endotheca located dorsal of tapering process, ending in subapical, backward-directed spines arranged in irregular collar; apical part subdivided and densely covered with minute microtrichiae giving dark colour.

Holotype: INDONESIA (Sumatra): Bukit Barisan Selatan National Park, Barisan Mts., Way Pitias , 800 m, 17–19.vi.2008, light trap [J. Oláh, Jr.] — 1 male (OPC, in alcohol).

Distribution: Indonesia (Sumatra).

Etymology: Complexa , referring to the uniquely complex dorsobasal processes of the gonopods.

Figures

FIGURES 8–11. Pseudoneureclipsis complexa, new species, holotype. 8—genitalia, left lateral; 9—genitalia, dorsal; 10—right gonocoxite, ventral; 11—phallic apparatus, left lateral.