Polymorphanisus scutellatus Banks, 1939

Uy, Christine Jewel C., Malicky, Hans & Bae, Yeon Jae, 2018, Review of the filter-feeding caddisfly subfamily Macronematinae (Trichoptera: Hydropsychidae) in tropical Southeast Asia, Raffles Bulletin of Zoology 66, pp. 664-703 : 693

Polymorphanisus scutellatus Banks, 1939

( Figs. 190, 191 , 200 , 220–222, 233)

Polymorphanisus scutellatus Banks, 1939: 55 [Holotype female; Borneo; MCZ].

Polymorphanisus scutellaris Banks ; Kimmins, 1955: 399 (incorrect spelling of scutellatus Banks ).

Diagnosis. Male. Antennae 50 mm in length. Antennal flagellum brown with darker striations. Forewing 20 mm and hind wing 14 mm in length ( Fig. 200 ). Mesoscutum without evident markings. Mesoscutellum with two oval markings as in Fig. 190 . Tibial spurs 1.3.2.

Male genitalia. Inferior appendages two-segmented, harpago as thick as coxopodite (Fig. 220). Phallus with wide opening in apex (Figs. 221, 222). Segment IX squarish, curving apicodorsad (Fig, 220). In dorsal view, segment X bifid with each half consisting of two lobes ( Fig. 233 ).

Female. Antennae 35 mm in length. Antennal flagellum same colouration as male. Forewing and hind wing length same as male. Mesoscutum without evident markings. Mesoscutellum with two larger dark spots covering almost whole area and in some individuals looking like lung shape ( Fig. 191 ). Tibial spurs 1.3.2.

Material examined. Indonesia: 2 males, 2 females, Aek Tarum, coll. H. Malicky, 6 March 1994 (HMPC) .

Distribution. Borneo, Sumatra, Java, Sulawesi.

Remarks. Barnard (1980) only observed one type of thoracic markings for this species ( Fig. 190 ), whereas Ulmer (1951) reported two variations. The specimens examined here exhibited two types of thoracic markings, and the males were different from the females ( Figs. 190, 191 ). The examined specimens possessed tibial spur formulas of 2.3.2, 1.3.3, or 1.3.3.

Figures

Figs. 183–192. Polymorphanisus thorax. 183, astictus, ocularis male and female thorax; 16, fuscus male thorax; 184–185, fuscus female thorax; 186, muluensis male and female thorax; 187, nigricornis male and female thorax; 188, quadripunctatus male and female thorax; 189, semperi male and female thorax; 190, scutellatus male thorax; 191, scutellatus female thorax; 192, unipunctus male and female thorax. Scale = 2 mm (189 redrawn from Barnard, 1980).

Figs. 193–201. Polymorphanisus male right fore- and hind wings. 193, astictus; 194, fuscus; 195, muluensis; 196, nigricornis; 197, ocularis; 198, quadripunctatus; 199, semperi; 200, scutellatus; 201, unipunctus. Scale = 5 mm (199 redrawn from Barnard, 1980).

Figs. 226–234. Polymorphanisus segment X of male genitalia. 226, astictus; 227, fuscus; 228, muluensis; 229, nigricornis; 230, ocularis; 231, quadripunctatus; 232, semperi; 233, scutellatus; 234, unipunctus. Scale = 0.5 mm (232 redrawn from Barnard, 1980).