Panscopus (Phymatinus) squamosus Pierce, 1913

Spanton, Timothy G. & Anderson, Robert S., 2016, A Taxonomic Revision of Weevils of the Genus Panscopus Schönherr (Coleoptera: Curculionidae: Entiminae: Tropiphorini), The Coleopterists Bulletin (mo 15) 70, pp. 1-86 : 35-36

Panscopus (Phymatinus) squamosus Pierce, 1913

( Figs. 12 , 38 , 61 , Map 13 )

Panscopus (Panscopidius) squamosus Pierce 1913: 394 . Leng 1920: 312.

Panscopus (Nocheles) squamosus ; Buchanan 1927: 28, 1936: 9; Schenkling and Marshall 1931: 40; Ting 1938; Blackwelder 1939: 66; Bousquet et al. 2013: 345.

Panscopus squamosus ; Hatch 1971: 289; O’ Brien and Wibmer 1982: 63; Bright and Bouchard 2008: 265.

Panscopus (Panscopidius) dentipes Pierce 1913: 395 . Leng 1920: 312.

Panscopus (Nocheles) dentipes ; Buchanan 1927: 28; Schenkling and Marshall 1931: 40.

Panscopus (Nocheles) squamosus dentipes ; Buchanan 1936: 9; Blackwelder 1939: 66.

Panscopus squamosus dentipes ; O’ Brien and Wibmer 1982: 63.

Type Material. Holotype, male: “Hilgard 1 Oreg.; Collection Wickham; Type l No.146401 U.S. N.M.; Panscopus squamosus Type Pierce” [Union County]. The holotype of P. dentipes , a male (Pullman, Washington, 13 April 94, J. W. Hungate collector; Type No. 14641 U.S. N.M.), was also examined.

Specific Epithet. The Latin adjective “ squamosus ” means scaly and is derived from the noun “ squama ” or scale.

Diagnosis. Specimens of this species are recognized by the long scape extending to the hind margin of the eye and females with four ventrites, in combination with the following of character states: elytral intervals even in elevation or at most slightly elevated near base; elytron with declivity in lateral view describing a continuous curve to apex; and setae more appressed than decumbent, varying from broad and scale-like to thin but flattened, and color cinereous.

Redescription. Females: Length 5.3–6.9 mm (n = 10); width across elytra 2.7–3.2 mm (n = 10). Males: Length 5.3–7.0 mm (n = 10). Color: Greybrown to dark brown, with irregular patches of grey scales and brown scales on elytra, integument, where visible through vestiture, rufotestaceous. Head: Transverse concavity at base of frons nearly absent; rostrum and frons apparently continuous in lateral view; rostrum and frons with shiny, thin, median longitudinal carina incomplete in some specimens, indistinct to absent in some specimens; lateral rostral carinae absent from most specimens, incompletely present in some; nasal plate indistinctly separated from dorsal surface of frons and rostrum, v-shaped carina present; vestiture of frons and rostrum of scattered decumbent to appressed setae, without clearly delineated clump of supraorbital setae. Antennae: Scape extended to or beyond hind margin of eye; scape vestiture of thin, apically directed decumbent to appressed setae, integument clearly visible; antennomeres 2–8 with long primary setae on distal portion of each article, and smaller, secondary setae scattered over surface, integument clearly visible. Pronotum: 1/5–1/6 wider than long, roundly convex on lateral margins, widest anterior to middle; surface coarsely rugose-tuberculate; median longitudinal sulcus thin, present length of pronotum; vestiture of scattered, decumbent to appressed, flattened setae. Elytra: Combined width approximately 2/3 length in females, males slightly narrower; nearly 50% wider than pronotum in females, slightly narrower in males; declivity, in lateral view, rounded, posterior margin not reflexed past vertical; declivity in females broadly rounded in lateral view, posterior margin reflexed beyond vertical; intervals subequally convex, alternate intervals not markedly raised above level of even intervals; all intervals with irregular rows of decumbent to appressed, flattened setae; scales of serial pits elongate, not round, about size of surrounding setae; elsewhere with vestiture of dense, overlapping, appressed, slightly elongate, striate scales. Legs: Tibiae 1 and 2 without obvious stiffened spines on ventral margin in most specimens; tibia 3 without obvious spines on ventral margin in most specimens; corbellar area of tibia 3 indistinctly closed. Abdomen: With 4 ventrites in females (ventrites 4 and 5 fused); 5 ventrites in males; ventrite 1 convex at middle in females, concave at middle in males. Genitalia: Females ( Figs. 12 , 38 ): Abdominal sternum VIII (spiculum ventrale), as in P. spantoni ; coxites 1 and 2 and styli as in P. spantoni ; vagina long, nearly twice length of coxites, membranous, with slightly crescent-shaped pair of sclerites anteriorly, near confluence of common oviduct; spermatheca broad and thick over most of its length. Males ( Fig. 61 ): Aedeagus decurved, with broad, spatulate projection apically, as in P. spantoni , manubrium of tegmen approximately as long as median lobe, without apical projection; median struts as long as median lobe including apical projection; internal sac with pair of sclerites apically, joined together in form of inverted v-shaped structure; membranous sac with field of sclerotized denticles distal to belt of tegmen.

Life History. Adults have been collected from February to August. Nothing else is known of the life history.

Geographical Distribution. This species lives in the Blue Mountains area of northeastern Oregon and eastern Washington, as well as in adjacent parts of western Idaho ( Map 13 ). In the former area, these weevils occur at elevations between 1,220 and 1,830 m .

Chorological Affinities. See this topic for P. spantoni above.

Material Examined. In addition to the holotype mentioned above, approximately 55 specimens were examined. USA: Idaho: Latah Co., Moscow , 31.III.1932, T.A. Brindley (1, UICM) . Valley Co., Cascade , 23.VI.1939 (1, USNM) ; Oregon: Baker Co., Wallowa-Whitman National Forest, McBride Campground, 9-12.VI.1989, E. Fuller, under rocks (1; CMNC) . Umatilla Co., Blue Mountains., Tollgate , 17.VII.1948, M.C. Lane (5, OSAC) ; Blue Mountains, Tollgate 1520 m, M.C. Lane (17, OSAC) ; Tollgate vic. Target Meadows 1520 m, 22.VII.1985, R.L. Westcott (1, ODAC) ; Tollgate, 8 km W., 1220 m, Roadside Meadow , 11.VII.1964, T. Schuh & J.D. Lattin, roadside meadow (1, UNSM) ; Blue Mountains , 11.VI.1940, K.M. Fender (2, USNM) ; Blue Mountains , 1830 m, 30.VIII.1923, M.C. Lane (4, USNM) ; Blue Mountains, Bone Springs 1770 m, 26.VI.1936, 11.VI.1940 (2, OSAC) ; Blue Mountains, Tollgate , 8.VI.1949, M.C. Lane (1, OSAC) ; Tollgate Road, Blue Mountains , 7.VI. 1938, Van Dyke collection (1, CASC) . Washington: Walla Walla Co., Walla Walla , 3.V.1940, M.C. Lane (2, OSAC) ; Blue Mountains, Godman Springs 1680 m, 3.VII.1940, M.C. Lane (5, OSAC) . Whitman Co., Pullman (2, USNM) ; Pullman , 26.IV. 1907 (1, WSU) ; Pullman , 10.IV.1956, V. Newhouse (1, WSU) ; Pullman , 5.IV.1913, M.A. Yothers (1, CASC) ; Pullman , 20.II.1945, May, C.V. Piper (3, USNM) . County unknown: Boulder Cave , 13.V. 1949, C.E. Horner (1, OSAC) . Assumed mislabeled: “Wyom. / Cle Ellum / 13.V.36” (1, FMNH) .

Figures

Map 13. Geographical distribution of Panscopus squamosus.

Figs. 9–16. Panscopus species, sternum 8 (spiculum ventrale), ventral (a) and left lateral (b) views. 9) P. acutisetus; 10) P. gemmatus; 11) P. spantoni; 12) P. squamosus; 13) P. michelbacheri; 14) P. johnsoni; 15) P. bakeri; 16) P. longus. Scale bar = 1 mm.

Figs. 37–40. Panscopus species, coxites and vagina, dorsal (a), ventral (b), and left lateral (c) views. 37) P. spantoni;

Figs. 59–61. Panscopus species, aedeagus, dorsal (a), ventral (b), and left lateral (c) views. 59) P. gemmatus; 60) P. spantoni; 61) P. squamosus. b = belt of tegmen; is = internal sac; mb = manubrium of tegmen; ml = median lobe; ms = median strut; t = tegmen. Scale bar = 1 mm.