Mecynotarsus pilbarensis, Kejval & Cz, 2013

Kejval, Zbyněk, 2013, Taxonomic revision of the Australian Notoxinae (Coleoptera: Anthicidae), Acta Entomologica Musei Nationalis Pragae (suppl.) 53, pp. 1-98 : 70-73

Mecynotarsus pilbarensis sp. nov.

( Figs 102, 103 , 182 , 206 )

Type locality. Australia, Western Australia, 1 km NNE of Millstream, 21°35′S 117°04′E.

Type material. HOLOTYPE: ♂, ‘ 1 km NNE Millstream, W.A. (21.35S 117.04E) 3–4.iv.1971, M.S. Upton [p] // ANIC Specimen [p, green label]’ ( ANIC). PARATYPES: 6 ♂♂ 2 ♀♀, same data as holotype ( ANIC) ; 4 ♂♂ 4 ♀♀, ‘3 Km. NWbyW. of Millstream HS., 21.34S 117.03E WA. 22 Apr. 1971 Upton & Mitchell [p] // AUST. NAT. INS. COLL. [p; green label]’ ( ANIC) ; 1 ♀, same data, except: 11.Apr. 1971 ( ANIC) ; 1 ♀, ‘ 5 km. SE. Of Millstream HS., 21.37S 117.06E WA. 17 Apr. 1971 Upton & Mitchell [p] // AUST. NAT. INS. COLL. [p; green label]’ ( ANIC) ; 2 ♂♂, ‘(21.35S 117.04E) Millstream, WA. Deep Reach, 8.xi.70, at light E.B. Britton [p] //ANIC Specimen [p; green label]’ ( ANIC) ; 1 ♀, ‘(21.35S 117.04E) Millstream, W.A. Crystal Pool, 28.x.70, at light E.B. Britton [p] // ANIC Specimen [p; green label]’ ( ANIC) ; 7♂♂ 3♀♀, ‘ 1km N of Millstream,W.A. (21.35S 117.04E) 9–10.iv.1971, M.S. Upton [p] // ANIC Specimen [p, green label]’ ( ANIC, 2 spec. ZKDC) ; 1♂ 1♀, ‘½km WNW Millstream , W.A.(21.35S 117.04E) 7.iv.1971, M.S. Upton [p] // ANIC Specimen [p, green label]’ ( ANIC, ZKDC) ; 4 ♀♀, ‘ 17 km. NbyE. Of Cane River HS., 21.56S 115.39E WA. 27 Apr. 1971 Upton & Mitchell [p] // ANIC Specimen [p; green label]’ ( ANIC) ; 4 ♂♂ 2 ♀♀, ‘ West Peawah R. WA 14.iv.1976 K. & E. Carnaby [p] // AUST. NAT. INS. COLL. [p; green label]’ ( ANIC) ; 1 ♀, ‘ 16.22S 125.12E W.A. Charnley Riv. 2 km SW Rolly Hill CALM Site 25/2 16-20 June 1988 I. D. Naumann [p] // at light, open forest near closed forest margin // Mecynotarsus sp. 6 det. T.A. Weir 1989 [p+h] // AUST. NAT. MUS. INS. COLL. [p; green label]’ ( ANIC) ; 1 ♀, same data, except: Mecynotarsus sp. 2 det. T.A. Weir 1989 [p+h]’ ( ANIC) ; 2 ♂♂ 5 ♀♀, same data, but lacking identification label by T. Weir ( ANIC, 2 spec. ZKDC) .

Description (holotype, male). Body length 2.1 mm. Body reddish to dark reddish brown, elytra with vague darker markings; legs and antennae reddish.

Antero-lateral margins of frons simple (not raised near antennal insertion). Gular rugules anteriorly coarser, contiguous to fused as in Figs 138, 139 . Clypeal granules minute. Setation of head rather coarsely hairy, comparatively long antero-medially on frons, mostly appressed, with some raised setae dorsally near base. Antennae moderately long; antennomeres III–V at most 1.8 times, X about as long as wide; setation mostly fine, coarser to scaly on basal four antennomeres.

Pronotum moderately transverse, 1.5 times as long as wide, its lateral margins rather strongly and somewhat unevenly convex in dorsal view; posterior collar narrow but distinct. Pronotal horn moderately long and wide, subtriangular, its posterior angles distinct in dorsal view; horn margins armed with 4 lobules on each side, apical lobule simple, broadly rounded; horn crest distinct, with coarse, partly contiguous rugules on margins, evenly lowering anteriorly; submarginal rugules rather distinct and mostly well-spaced; median rugules partly fused and forming irregular sculpture, with several minute granules scattered posteriorly. Setation whitish (laterally) to reddish (dorsally), scaly, much finer / sparser to indistinct on pronotal horn dorsally; scales on pronotal disc of two sizes, generally appressed and very densely spaced, larger scales widely rounded apically, slightly glossy; antebasal paired setae present and rather long laterally, absent medially, several additional tactile setae present antero-laterally.

Elytra 1.6 times as long as wide; omoplates and postbasal impression absent. Setation reddish and brownish, forming rather sharply outlined brownish markings ( Fig. 206 ), scaly and appressed, evenly ordered; scales shortly oval, slightly glossy, very dense and entirely covering surface ( Fig. 182 ); erect tactile setae absent.

Male characters. Sternum VII nearly simple. Tergum VIII and aedeagus as in Figs 102, 103 .

Variation. Body length (♂ ♀) 2.0– 2.4 mm. Pronotal horn with 3–4 lobules on each side; median rugules separate to fused and forming irregular sculpture. Brownish markings of elytra may be both conspicuous, mostly so ( Fig. 206 ), and vaguely indicated to absent.

Differential diagnosis. Mecynotarsus pilbarensis sp. nov. resembles M. exophthalmus sp. nov., M. kingii , and M. magelae sp. nov. in external characters (body form, setation). It differs from M. exophthalmus sp. nov. mainly by the evenly shaped vertex (lacking narrow impressions along the median margins of the eyes), from M. kingii by the short frontal portion of the head with distinctly narrowing, simple margins, and from M. magelae sp. nov. by the wider and longer horn crest with separate rugules on the margins (never fused and forming evenly shaped rim), the numerous median rugules that are sometimes fused and forming an irregular sculpture (single, rather evenly shaped, longitudinal median rugule in M. magelae sp. nov.), and by the sparser fringe of pale, narrower scales on the posterior collar of the pronotum (dense fringe of whitish, wider scales in M. magelae sp. nov.). Moreover, males of M. pilbarensis sp. nov. can be easily distinguished from those of the other mentioned species by the shape of the parameres (cf. Fig. 103 versus Figs 48 , 68 , 75 ).

Etymology. The species name is a Latin adjective pilbarensis , - is, - e; named in reference to the type locality, which is located in the Pilbara region in Western Australia.

Distribution. Australia: Western Australia.

Figures

Figs 98–103.98–101 – Mecynotarsus pilicornis sp. nov.: 98 – tergum VIII, 99 – tegmen in dorsal view, 100 – median lobe in dorsal view, 101 – aedeagus in dorsal view (Gayndah). 102–103 – M. pilbarensis sp. nov.: 102 – tergum VIII, 103 – aedeagus in dorsal view. Scale (0.5 mm):A – Fig. 101; B – Figs 98–100, 103, 102.

Figs 177–185. Setation of elytra: 177 – Mecynotarsus kingii MacLeay, 1872 (Shoalhaven River), 178 – M. nobilis sp. nov., 179 – M. obesus sp. nov., 180 – M. parvulus sp. nov., 181 – M. phanophilus sp. nov., 182 – M. pilbarensis sp. nov., 183 – M. regalis sp. nov., 184 – M. speciosus sp. nov., 185 – M. weiri sp. nov.

Figs 202–209. Habitus: 202 – Mecynotarsus magelae sp. nov. (Charnley River), 203 – M. mastersii MacLeay, 1872 (Cedar Creek National Park), 204 – M. obesus sp. nov., 205 – M. parvulus sp. nov., 206 – M. pilbarensis sp. nov. (paratype, Millstream Homestead), 207 – M. regalis sp. nov., 208 – M. weiri sp. nov., 209 – M. ziczac King, 1869 (Newcastle Grain Terminal).

Figs 133–140. 133 – Mecynotarsus setulosus sp. nov., head in lateral view (paratype, Fowlers Gap), 134 – M. excavatus sp. nov., apex of elytra in female (paratype, Aoue-Enga), 135 – M. grandior sp. nov., apex of elytron in male (paratype, Kimberley Research Station), 136 – M. obesus sp. nov., metatibia with tarsus (paratype, Hull River), 137 – M. excavatus sp. nov., male protibia (paratype, Aoue-Enga). 138–140 – head in ventral view: 138 – M. auripilosus sp. nov. (paratype, Kambah Pool), 139 – M. bullatus sp. nov. (paratype, Regans Ford), 140 – M. ziczac King, 1869 (Swain Reefs).

Figs 47–52. 47–48 – Mecynotarsus exophthalmus sp. nov.: 47 – tergum VIII, 48 – aedeagus in dorsal view. 49–50 – M. fallax sp. nov.: 49 – tergum VIII, 50 – aedeagus in dorsal view. 51–52 – M. festivus sp. nov.: 51 – tergum VIII, 52 – aedeagus in dorsal view. Scale (0.1 mm): A – Fig. 50; (0.5 mm): B – Fig. 51; C – Figs 47–49, 52.

Figs 67–73. 67–68 – Mecynotarsus kingii MacLeay, 1872: 67 – tergum VIII, 68 – aedeagus in dorsal view. 69–71 – M. kreusleri King, 1869: 69 – sternum VII, 70 – tergum VIII, 71 – aedeagus in dorsal view. 72–73 – M. leai Pic, 1942: 72 – tergum VIII, 73 – aedeagus in dorsal view. Scale (0.5 mm): A – Figs 70–73; B – Fig. 69; C – Fig. 67; D – Fig. 68.

Figs 74–81. 74–75 – Mecynotarsus magelae sp. nov.: 74 – tergum VIII, 75 – aedeagus in dorsal view. 76–79 – M. mastersii MacLeay, 1872: 76 – sternum VII, 77 – tergum VIII, 78 – tegmen in dorsal view, 79 – median lobe in dorsal view. 80–81 – M. mollis sp. nov.: 80 – tergum VIII, 81 – aedeagus in dorsal view. Scale (0.2 mm): A – Fig. 75; (0.5 mm): B – Fig. 76; C – Fig. 77; D – Figs 80, 81; E – Figs 74, 78, 79.