Galenidae Alcock, 1898

Khosravi, Erfan, Sari, Alireza, Mirziee-Ataabadi, Majid, Gholamalian, Hossein & Hyžný, Matúš, 2022, A new species of Galene de Haan, 1833 (Galenidae: Brachyura) from the middle Miocene of Zagros Mountains, Iran, Zootaxa 5124 (2), pp. 139-154 : 144

Family Galenidae Alcock, 1898

Remarks. Galene has been classified in the Xanthidae (e.g., Balss 1957; Glaessner 1969; Takeda 1976), whereas Guinot (1969a, 1969b, 1969c, 1971) assigned Galene to “ Goneplacidae pilumniens sensu lato ”. This suggestion was followed by a number of scholars ( Ng 1998; Schweitzer 2000; Ng e t al. 2001; Hsueh & Huang 2002; Ng et al. 2008), who classified Galene within (or close to) the Pilumnidae . Ng et al. (2008) summarized the taxonomic history of Galenidae , which was included in the superfamily Pilumnoidea .

The family Galenidae Alcock, 1898 , as currently recognized, consists of four genera ( Ng et al. 2008; De Grave et al. 2009; Ng & Guinot 2021; Ng & Mitra 2021): Dentoxanthus Stephensen, 1946 ; Galene de Haan, 1833 ; Halimede de Hann, 1835 ; and Parapanope de Man, 1895 . The fossil record of Galenidae is limited to representatives of Galene and Halimede ( Schweitzer et al. 2010) .

Although there are some difficulties in a detailed morphological diagnosis of Galenidae, Guinot et al. (2013) denote the emergence of the penis from P5 coxo-sternal condyle as a diagnostic trait of this family; however, it is not usually possible to study such fragile elements in fossils.