Eupograpta kottae, Raven, Robert J., 2009

Raven, Robert J., 2009, Revisions of Australian ground-hunting spiders: IV. The spider subfamily Diaprograptinae subfam. nov. (Araneomorphae: Miturgidae), Zootaxa 2035, pp. 1-40 : 17-20

Eupograpta kottae sp. nov.

( Figs 10 a–c, 11a–c, 12a, b, 25a)

Material examined. Holotype. Male, Durokoppin Nature Reserve 31°30'S 117°44'E, Western Australia, 3–14 November 1988, D. Mitchell et al., DKR 83 ( WAM 97/243).

Paratypes. Allotype female, Boorabbin, 3115' S 12004 'E, Western Australia, July 1980, W. F. Humphreys et al., WA Goldfields survey, BNR 1, isolate Callitris heath ( WAM 97/189). Western Australia: 2 males, Perth Airport, 3158'03" S 11558 '11"E, 10 May–24 June 1993, wet pitfalls (PA 5), M. S. Harvey, J. M. Waldock ( WAM 98/1676, 1677); male, Talbot Rd Reserve, 3152'24" S 11602 '52"E, 24 September –18 November 1993, wet pitfalls ( TR 3), J. M. Waldock et al. ( WAM 98/1680); female, same data but 28 July –23 September 1993, J. M. Waldock et al. ( WAM 98/1682); 3 females, same data but 18 November–10 December 1993, J. M. Waldock ( WAM 98/1683–5); 2 males [1j.], same data but 24 June –28 July 1993, M. S. Harvey, J. M. Waldock ( WAM 98/ 1689–91); female, same data but dry pitfalls, 22–29 August 1993, J. Dell ( WAM 98/ 1692); 2 males, Talbot Rd Reserve ( TR 4), 3152'23" S 11602 '46"E, 24 September–18 November 1993, J. M. Waldock et al. ( WAM T74023, T74024); 1 female, same data but 23 November–5 December 1993, J. Dell ( WAM 98/1744); 1 male, Talbot Rd Reserve ( TR 1), 3152'05" S 11603 '04"E, 24 September–18 November 1993, J. Waldock et al. ( WAM T 74026); 1 female, Red Hill, Toodyay Rd, top of Darling Scarp, 3151' S 11606 'E, 6 August 1996, B. Y. Main ( WAM 98/1686); 1 male, Bold Park (BP3), wet pitfall, 3156'33" S 11546 '13"E, 20 May–20 July 1993, M. S. Harvey and J. M. Waldock ( WAM 98/1687); 1 male, Bold Park (BP5), 3157'14" S 11546 '16"E, 20 May–20 July 1993, M. S. Harvey and J. M. Waldock ( WAM T74019); 1 male, Hartfield Park, Forestfield (HF1), wet pitfalls, 3200'00" S 11559 '43"E, 16 April–19 June 1996, J. M. Waldock, P. West, A. Longbottom ( WAM T74025).

Diagnosis. Females differs from those of E. anhat sp. nov. in that the median septum of the epigyne ( Fig. 12 a) is widest distally.

Etymology. Named for Dr Patricia Kott, ascidian taxonomist, Queensland Museum, originally from Western Australia, in recognition of her tireless efforts to raise the profile of taxonomy and the need for more, funded taxonomic positions in Australia.

Description. Male (holotype WAM 97/243)

Carapace 3.00 long, 2.31 wide. Abdomen 3.56 long, 2.19 wide. (Cephalothorax slightly collapsed and abdomen folded under at 90.) Colour. Carapace fawn with black stripes from each PLE down carapace; broken black areas along margins of carapace; legs not banded. Abdomen ( Fig. 11 b) dorsally pallid with two long medial stripes joining just past midpoint, two stripes lateral of that; ventrally pallid. Small stripe down each chelicera. Carapace. With gradual anterior constriction. AME set on anterior clypeus. Fovea short, at ca. midpoint of carapace; few long anteromedial bristles, 3–5 between MOQ, 2–3 lines of 4–6 long curved bristles anterior to AME on clypeus. Clypeus about twice AME diameter, convex, steep. Fovea moderately long. Short brown bristles uniformly over carapace. Carapace ridge flat between back margin and PME. Clypeus anterior to AME and ALE tumid, pallid. Eyes. PME largest. Two recurved rows, posterior more so. Eyes of anterior row set close together; in anterior view, smaller AME set above centres of ALE. Viewing directions: AME up, front, side; ALE front, side; PME up, PLE to side and slightly to back. All eyes encircled with dark shadows. Chelicerae. P1 small basal and 2 larger, R2 large basal. Small triangular boss. Labium and maxillae slightly collapsed, see female. Sternum. Cordate with uniform cover of long bristles; low radial ridge/mound opposite base of each coxae. Legs. Trochanters all deeply notched. No tibial fracture, no tarsal rod. Pilosity: broad long brown bristles of fine hairs. Distinct long bristles off apical patellae. I: 2.77, 1.23, 2.46, 2.15, 1.69, 10.30. II: 2.77, 1.31, 2.23, 1.92, 1.54, 9.77. III: 2.46, 1.15, 2.00, 1.85, 1.38, 8.84. IV: 3.31. 1.31, 3.15, 3.0. Palp: 1.23, 0.69, 0.54, –, 1.54, 4.00. Trichobothria. On tibiae, 2 rows 6–8 for length; metatarsi, irregular rows of 8–10; tarsi, 1 row of 8–10, subapical trichobothria longest. Spines. 2 basal spines on dorsal femora very long. I: fe p2d2r1; pa0; ti p 2v 2.2.2; me v2.2. II: fe p3d3r3; pa0; ti p2r 1v 2.2.2; me p 1v 2.2, me spines very long. III: fe p4d3r3; pa0; ti p2d2r 2v 2.2.2; me p4r 4v 2.2.1. IV: fe p4d3r2; pa0; ti p2d2.1r 2v 2.2.2 me p3+2l r 7 v 2.2.1. Palp: fe p1d3r2; pa p1d1; ti p1d1. Scopula . Long hair-like setae, thin but distinct for distal two-thirds of metatarsi I, for distal half of metatarsi II–IV. Long hair-like setae, moderately dense, entire on tarsi I–IV almost obscures cuticle on I. Claw tufts small, dense, no division evident. Claws. Moderately long, curved with 6–8 long teeth evident above tufts. Spinnerets. ALS as long as PLS but ALS thicker. PLS with slightly longer apical segment. Palp ( Figs 10 , 11 a). Patellae squat, dorsal projection a sclerotised thumb, opposed to sclerotised back dorsal tibial edge. Tibia barrel-shaped, practically only an apophysis, "sandwich" with unsclerotised midzone. RTA a very long, elaborate, twisted process with ental grooves and digitiform process twisting apically into fluted cylindrical tip. Long triangular process on proventral corner and collar ventrally. Distal two-thirds ventrally glabrous, saddle-shaped. Cymbium dorsally with triangular process basally against one on tibia; retrobasal sharp corner with excavation more basally; retrolateral cymbium excavate with long groove and distal process; groove on thick ridge and length of groove retromargin with dense line of short black bristles as comb; promargin of cymbium with long densely padded pallid hairs (removed for SEM). Embolus origin long, ovoid; anticlockwise to narrow tip above embolus origin. A large flat median apophysis with edges curved up and returns with small tooth and rounded flange; beside that a sclerotised scooped, boat-shaped conductor surrounding embolus. Massive tegulum. No scopula dorsally on cymbium.

Female (allotype WAM97/189)

Carapace 3.45 long, 2.60 wide. Abdomen 4.70 long, 2.85 wide. Like male except: Colour. Carapace yellow brown with separated dark semicircles along margin band and broad dark band from PLE to back; chelicerae red-brown with dark vertical medial band. Abdomen dorsally pallid; centrally for anterior half with dark "shoulders" rising to just over the dorsal surface and medial of them two narrow dark lines anteriorly well apart, diverging slightly and shortly but converging gradually (i. e., forming a long narrow “V) to join just behind midpoint, irregular mottling posterior of that forming pallid area centrally, laterally with short dark broken longitudinal diagonal lines and ventrally with vaguely discernible broad pallid ”V with scattered irregular mottling over it. Sternum and legs yellow to orange brown without dark markings or bands. Carapace. Light uniform pile of large dark hairs centrally directed; midline from fovea to PME with a wide band of long erect bristles and ca. 6 long bristles directed horizontally forward from clypeus. Fovea distinct, crosses into posterior declivity. Eyes. In two rows Chelicerae. 2R, 3P. Legs. Trochanteral notches broad but deep, distinct. RCH distinct, moderately large. I: 2.10, 1.40, 1.95, 1.70, 1.05, 8.20. II: 2.35, 1.40, 1.90, 1.50, 1.15, 8.30. III: 2.60, 1.25, 1.65, 1.45, 1.00, 7.95. IV: 3.05, 1.40, 2.75, 3.45, 1.15, 11.80. Palp: 0.90, 0.50, 0.65, –, 1.05, 3.10. Scopula as for D. striola . Claws. I–IV with ca. 4–5 teeth, becoming smaller toward base; tufts on IV noticeably smaller than on I–III. Spines. Leg patellae without spines. I: fe p1d2; ti v 2.2w long; me v2 very long, basal. II: fe p2d2; ti v1.1; me v2 long. III: fe p3d3r2; ti p2r 2v 2.2.2; me p1r 1v 2.1. IV: fe p1d3r1; ti p2d2r 2v 2.2.2; me p1.2.2r2.2.2v2.1. Palp: fe d1.2; pa p1d1 + apical 1; ti p3d2; ta p3d1r 3 v 2 distal. Spinnerets. As in D. striola with wide colulus. Epigyne. Large, two strongly sclerotised lateral ridges bracket broad hirsute medial scape narrowing strongly in posterior third to form subdistal knob, in distal third scape edges form ridge below which is small slit-like invagination then expanded slightly sclerotised fold below which is a large unsclerotised concavity.

Distribution and habitat ( Fig. 25 a). Known only from forests on the margins of Perth, southwestern Western Australia and at Boorabbin in the Goldfields to the west.

Figures

FIGURE 10. Eupograpta kottae sp. nov., male, palpal tibia, cymbium and bulb and tarsus: a, bulb, ventral view; b, c, median apophysis ventral views, distal edge (b) and proventral edge (c); d, e, RTA, retrolateral (d) and ventral (e) view; f, end of tarsus showing claws and tufts, lateral view; g, bothrium base.

FIGURE 11. Eupograpta kottae sp. nov., holotype male (a) and allotype female (b, c): a, patella to cymbium, ventral view; b, c, abdomen, dorsal (b) and ventral (c) view.

FIGURE 12. Eupograpta kottae sp. nov., allotype, female: a, b, epigyne, external (a) and internal (b) views.

FIGURE 25. Distribution records of Miturgidae in Australia: a, Diaprograpta, Eupograpta, Mitzoruga and Nuliodon; b, all records of Miturgidae for Australia.