Eratigena hispanica Bolzern, Angelo, Burckhardt, Daniel & Hänggi, Ambros, 2013

Bolzern, Angelo, Burckhardt, Daniel & Hänggi, Ambros, 2013, Phylogeny and taxonomy of European funnel-web spiders of the Tegenaria-Malthonica complex (Araneae: Agelenidae) based upon morphological and molecular data, Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 168 (4), pp. 723-848 : 767-768

ERATIGENA HISPANICA ( FAGE, 1931) COMB. NOV.

( FIG. 12F–G, L–M )

Tegenaria hispanica Fage, 1931: 212 , 213, fig. 48, only female; Ribera, 1978: 30–32, figs 1–5, male.

Pseudotegenaria hispanica: Lehtinen, 1967: 261 .

Types

Holotype and paratype. Spain: Catalonia: Tarragona, Tortosa, Cova d’en Rubi , 2 ♀ ( MNHN, 1979, 787), 20.v.1914, Fage, 1931 .

Other material examined

Spain (1 ♂, 2 ♀) .

Diagnosis

See the Diagnosis section for E. herculea .

Description

Measurements: Male (N = 1): CL 4.34, CW 3.14, STL 2.14, STW 2.02, OL 2.97, OW 1.65. Leg I (7.33, 1.71, 7.43, 7.71, 3.61), II (6.32, 1.73, 5.78, 7.17, 3.1), III (5.53, 1.36, 4.89, 6.93, 2.86), IV (6.6, 1.43, 6.31, 9.51, 3.64). Pedipalp (2.31, 0.76, 1.34, 1.81), bulbL 0.66. Female (N = 1): CL 5.32, CW 3.54, STL 2.43, STW 2.16, OL 5.01, OW 2.7. Leg I (7.42, 1.98, 7.58, 7.99, 3.53), II (6.77, 1.93, 6.27, 7.56, 3.59), III (6.39, 1.78, 5.73, 7.89, 3.2), IV (7.65, 1.98, 7.51, 10.68, 3.92). Pedipalp (2.74, 0.96, 1.97, 2.84). EPL 0.49, EPW 0.76, ATL 0.33, ATW 0.28. Eyes (somewhat reduced): PME 0.13, PLE 0.13–0.17, AME 0.10–0.11, ALE 0.16–0.17. Eye distances: PME- PME 1–1.5 x PME, PME–AME 1 x PME, PME–PLE 1.5–2 x PME, PME–ALE 1.5–2 x PME, AME–AME 1–1.5 x AME, AME- ALE 0.5–1 x AME, CLY1 3–3.5 x AME, CLY2 1.5–2 x ALE.

Male palp: RTA with two branches, lateral branch only moderately protruding, lobe-like dorsal branch strongly sclerotized and protruding, distally with short, claw-like appendix. Short dorsal spike on male palp tibia absent. Embolus length 1.75–2 x CB, originating at 7–8 o’clock position, distal tip at 4 o’clock position (bulb in Fig. 12L–M twisted). Conductor with distal portion moderately elongated, as long as wide, reaching distal margin of alveolus, lateral margin folded. Terminal end simple, long, drawn out, and pointed. Transversal ridge of conductor membranous, lamelliform. Conductor membranously connected to tegulum. MA originating at 5 o’clock position, protruding, longer than wide, distally with spoon-like sclerite. MA membranously connected to tegulum.

Epigyne and vulva: Epigyne with distinct posterior sclerite, forming strongly sclerotized, triangularly to rectangularly shaped, and protruding pocket, opening posteriad. Epigynal teeth present, originating distally of posterior sclerite, pointing posteriomediad. CO located anteriolaterally of posterior sclerite at a rectangularly shaped membranous area. Vulva consists of distinguishable CD, RC, and FD. CD short, with long, distally strongly convoluted appendix. RC irregularly formed and sclerotized with several ‘chambers’, separated by two x the diameter. FD only represented by small, leaf-shaped appendages.

Other important characters: Cheliceral promargin with three, retromargin with nine to ten teeth. Colulus rectangularly shaped with distal margin more or less straight or moderately w-shaped. Distal segment of PLS longer than basal segment. PMS with one prominent minor ampullate gland spigot and three to four cylindrical gland spigots laterally. Tarsal trichobothria on cymbium and palp tarsus absent. Tarsal trichobothria eight to nine. Small teeth on paired claws of leg I 13–14. Leg spination: male palp (2–1–1–0, 2–0–0, 1–1+1p–0–0), female palp (2–1–0–0 or 2–1–1–0, 2–0–0, 2–2p–0–0), leg femora (2–3–3–0 or 2–3–4–0 or 2–4–3–0, 2–3–4–0, 2–2–4–0 or 2–3–4–0, 2–2–2–0 or 2–3–2–0 or 2–3–3–0), patellae (all 2–0–0), tibiae [2–0–0–4p or 2–1–0–4p or 2–2–0–4p (lateral spines only in female), 2–2–0–4p, 2–2–2–1+3p, 2–2– 2–1+3p or 2–2–2–4p], metatarsi (0–0–0–4p+1, 0–2–0– 4p+1 or 0–2–1–4p+1, 0–5–4–4p+1 or 1–5–4–4p+1, 0–5–4–4p+1 or 1–5–4–4p+1), tarsi (I & II 0, III 0 or 0–0–1–0 or 0–1–2–0, IV 0–2–2–0 or 0–2–3–0).

Coloration: Carapace not darkened (troglobiont?). Sternum with indistinct pale median region. Opisthosoma brown-yellowish, only cardiac mark darker. Legs without a pattern.

Distribution

Reported from Catalonia and Valencia ( Ribera, 1978; Ribera & Barrientos, 1986).

Figures

Figure 12. A, B, Eratigena herculea; C, D, H, N, O, Eratigena arganoi; E, K, Eratigena sicana; F, G, L, M, Eratigena hispanica; I, J, P, Q, Eratigena sardoa; R–T, Eratigena picta. Female epigyne in ventral (A, C, F, I) and vulva in dorsal view (B, D, E, G, J); male left palp in ventral (L, N, P, R) and retrolateral views (M, O, Q, S); frontal face of male E. arganoi (H); habitus of female E. sicana (K); habitus of two males of E. picta (T). Scale bars = 0.5 mm.