Cyamops femobrunneus, Grimaldi, 2009

Grimaldi, David A., 2009, The Asteioinea of Fiji (Insecta: Diptera: Periscelididae, Asteiidae, Xenasteiidae), American Museum Novitates 3671, pp. 1-60 : 23-26

Cyamops femobrunneus , new species

Figure 14

DIAGNOSIS: Face with peristomal setae and well-developed vertical ridge; arista without bifurcate branches or at best with 3 minute ones near bases of proximal branches; femora almost entirely light brown; forefemur without ctenidium, mesocoxa without small comb of thick setae on posterior margin; male terminalia with left surstylus long, curved, total length 4.8X greatest width.

DESCRIPTION: ThL 5 1.81 mm (holotype). Body coloration: Head mostly dark brown, velvety black within and immediately around ocellar triangle, ocelli light; velvety brown in middle of frons; dorsal portion of frontoorbital plates lighter, shiny brown, anterior portion of FOPs with silvery microtomentum. Antenna with scape, pedicel, basal flagellomere yellowish. Face yellowish, with slight silvery microtomentum beneath antennae, thick microtomentumon narrow strips of cheeks beneath eyes; middle of face with light bluish microtomentum. Eyes pink. Clypeus dark brown, mouthparts very light yellow. Postocciput dark brown. Thorax entirely dark brown (scutum, scutellum, pleura). Pleura with very slight microtomentum. Forecoxae light, anterior surface with thick silvery microtomentum; other coxae light brown. All femora brown, with proximal end lighter; tibiae and tarsi yellowish, distal tarsomere dark brown. Halter light brown; wings hyaline (no patterns), very slightly infuscate. Abdominal tergites lighter brown than scutum and scutellum, except epandrium and tVII (which are darker); sternites light brown.

Head: Frons with large pair of proclinate orbital setae, lying immediately lateral to pair of reclinates, which are slightly shorter. Anterior to reclinate is row of minute ca. 6 reclinate setulae. No ocellar, postocellar, or inner vertical setae present; outer verticals large, lateroclinate; postocellars present as minute pair of setulae. Eye with large facets on anterior surface, at margins where face is pinched; these facets ca. 2X diameter of other facets. Eyes bare, no setulae between facets. Face very narrow between eye margins, width less than 2X facet diameter; face with narrow, shallow, vertical carina that is polished brown to yellow. Pair of pseudovibrissae just ventral to narrowest portion of face, pointed dorsad, fine, lengths ca. 4X diameter of largest eye facets. Row of approximately 8 long parafacial setae (alterating with minute setula), 4 pairs on face. Antenna with pedicel cuplike, ventral portion of rim with several long setae; basal flagellomere projecting forward (not downward). Arista without small basal article, plumose, with 7 long dorsal branches and 2 shorter ventral branches (towards apex) (not including apical fork), ca. 7 short mesal branches; dorsal branches either without small secondary branches (though 2–3 branches have slightly thickened areas in middle and are slightly bent, suggesting loss of secondary branches) or there are at most 2 minute secondary branches on most proximal branches of arista. Cheek shallow, with ca. 4 setae; postgena with 3 stiff setae on each side. Clypeus very narrow, greatest width ca. 1/3 that of oral cavity. Palpi small; labellum with fringe of fine, light setae on lateral margins.

Thorax: Scutum with 2 short rows of long acrostichals, ca. 4 acrostichals per row. Three pairs strong dorsocentrals; anterior pair shortest, posterior pair longest; ca. 5 short, fine setae in row in front of anterior dc. Scutellum with two pairs of scutellar setae, posterior pair longer, ca. 1.3X length of anterior pair. Pleura with 2 thick, long notopleural setae, approximately equal in size; katepisternum with 2 setae, 1 thick and long, the other fine and ca. 0.5X the length. Thorax deepest at level of katepisternum and mesocoxa. Venter of thorax broad, with fore coxae well separated and mesosternite very broad. All setae on venter of thorax fine, none especially thick and stiff. Legs with few distinctive setae, except thick, black, long seta at apex of midtibia. Tarsomeres expanded in length distally; distal tarsomeres broad, flat, claws large, pulvilli large. Wing: Calypter with fringe of long, fine setae; vein C extends to apex of vein M, without humeral break and with faint Sc break; Sc long, seems to fuse apically with R 1; vein R 2+3 very long, extended well past level of cross vein dm-cu, otherwise venation typical of Cyamops .

Abdomen: Short, length only slightly longer than that of thorax, slightly more narrow than thorax. Tergites and sclerites well developed, pleural membrane narrow. Epandrium heavily sclerotized, capsulelike, symmetrical, without long ventral lobes. Tergite VII asymmetrical, with apex (ventral to epandrium) blunt and sclerotized. Surstyli highly asymmetrical, left one long (length ca. 4.8X greatest width) and strongly curved; right one short and thick (as figured).

TYPES: Holotype, male: Taveuni : Cakaudrove Prov., 5.6 km SE Tavuki Vlg, Devo Peak 1187 m, 24–31.X.2002, Malaise 1, coll. E. Schlinger and M. Tokota’a, 16.843 ° S, 179.965 ° W, FBA 160268; paratypes, 4 males, same data, nos. FBA 020111, 160271, 149933 (dissected, no. 64), 160273. In BPBM and AMNH.

ETYMOLOGY: From femur and brunnea (L., ‘‘brown’’), in reference to the entirely brown femora.

Figures

Fig. 14. Structures of Fiji Cyamops spp. A. Male terminalia (posterior view) of Cyamops femobrunneus. B, C. Aristae of C. femobrunneus (B) and C. femoctenidius (C).