Crettaros santibanezi, Cruz-López & Francke, 2015

Cruz-López, Jesús A. & Francke, Oscar F., 2015, Cladistic analysis and taxonomic revision of the genus Karos Goodnight & Goodnight, 1944 (Opiliones, Laniatores, Stygnopsidae), Zoological Journal of the Linnean Society 175 (4), pp. 827-891 : 867

CRETTAROS SANTIBANEZI SP. NOV.

( FIGS 41 , 42 , 57 , 61A , 63D )

Type material: MEXICO: Querétaro: ♂ holotype , 1 ♂ and 1 ♀ paratypes [ CNAN-T0089 and CNAN-T0090 (15.xi.2009; O. Francke, A. Valdez, C. Santibáñez, and

J. Cruz)] (21°7′32.7″N, 99°38′21.04″W). Municipio Pinal de Amoles, 200 m from intersection Pinal de Amoles-Bucarelli.

Material examined: 1 ♂ and 1 ♀ [ CNAN (17.vi.2007; L. Pastrana)] (21°8′0.88″N, 99°27′29.05″W). Municipio Jalpan de Serra , La Yerbabuena . 2 ♀ [ CNAN (17.vi.2007; L. Pastrana)] (21°17′15″N, 99°8′16″W). Municipio Landa de Matamoros , El Madroño .

Etymology: Patronym, dedicated to Carlos Santibáñez- López, friend and colleague, who has contributed to the knowledge of scorpions in Mexico.

Diagnosis: Epigean species. Dorsum covered by small, rounded tubercles. On the male penis the two basalmost pairs of lateral setae are separated ( Fig. 42B ).

Description: Male holotype: Measurements: scutum length 2.5, maximum scutum width 2.1. Legs. I 1.25/ 0.50/0.85/2.20, II 1.25/0.55/0.85/1.10, III 1.35/0.60/1.15/ 1.30, IV 1.80/0.65/1.70/1.90. Dorsum. Covered almost completely by small, rounded tubercles, mesotergal area I generally with two central pairs of tubercles. Opisthosoma slightly convex in lateral view. Mesotergal areas II and III with a median hump, without armature ( Fig. 63D ). Ocularium slightly separated from anterior margin of prosoma, low, slightly elevated above the eyes, blunt, covered by small tubercles ( Fig. 41F ). Lateral clear areas projected in small, teardropshaped projections, inconspicuous. Lateral clear areas on the apices of area V and free tergites I–III, rounded, not protruding significantly. Free tergites with median row of small tubercles. Venter. Coxae I and II with a median row of rounded tubercles, coxa III covered completely by small tubercles, coxa IV covered by tubercles and numerous setae, with a small dorsoectal spiniform tooth. Stigmatic area shaped like an invert- ed ‘Y’, with lateral margins straight, parallel ( Fig. 41D ). Pedipalps. Patella with one mesal spiniform tubercle. Legs. Very similar in ornamentation and size, covered by numerous small tubercles and setae. Trochanter IV with numerous tubercles, ventral ornamentation of femur IV slightly bigger. Femur IV curved, constrained basally, slightly expanded in the middle.

Figures

Figure 41. Crettaros santibanezi sp. nov. A, habitus, dorsal view. B, legs IV, mesal view. C, habitus, lateral view. D, habitus, ventral view. E, scutum, dorsal view. F, ocularium, frontal view. Scale bars: A = 2.5 mm, B, D, and E = 1.5 mm, C = 1 mm, F = 0.5 mm. The dark lines on (D) indicate the stigmatic area.

Figure 42. Crettaros santibanezi sp. nov. male genitalia. A, dorsal view. B, ventral view. C, lateral view. White arrows indicate ventral microsetae in (B) and parastylar setae in (C).

Figure 57. Distribution map of the species of the Karos genus-group. Black triangles, Monterella tuberculata; black square, Mictlana inops comb. nov. (only locality of material examined was mapped); black circles, Crettaros santibanezi sp. nov.; white circles with black outline, Crettaros valdezi sp. nov.; black crosses, Chapulobunus unispinosus; inverted black triangles, Karos parvus.

Figure 61. A, detail of one of the lateral clear areas forming tubercles on scutum of Crettaros santibanezi sp. nov. B, detail of one of the lateral clear areas projected in tubercles on scutum of Karos singularis sp. nov. C, detail of one of the lateral clear areas projected in tubercles on scutum of Karos tersum sp. nov. D, detail of one of the lateral clear areas projected in tubercles on scutum of Huasteca silhavyi sp. nov. E, detail of one of the lateral clear areas projected in tubercles on scutum of Montabunus foliorum. F, detail of one of the lateral clear areas projected in tubercles on scutum of Monterella tuberculata. G, extra row of pegs in Chapulobunus poblano sp. nov., indicated by arrows. H, dorsal apophyses on trochanter II in male of Karos singularis sp. nov. I, detail of mesotergal areas III and IV in Montabunus foliorum; dotted lines indicate the sulcus between these areas. J, detail of mesotergal areas III and IV on Chapulobunus unispinosus; dotted lines indicate the sulcus between these areas. These images are not at the same scale.

Figure 63. A, dorsal view of the anal bulge on Chapulobunus poblano sp. nov.; arrow indicates the bulge. B, lateral view of Huasteca silhavyi sp. nov., line indicates the very high lateral channel area. C, laterodorsal view of scutum of Crettaros valdezi sp. nov., showing the median humps on areas II and III indicated by dotted circles. D, posterodorsal view of scutum of Crettaros santibanezi sp. nov., showing the median humps on areas II and III indicated by dotted circles. E, dorsal view of trochanter IV of Karos barbarikos; arrow indicates the tridentate apophysis. F, anterodorsal view of scutum of Karos barbarikos, the black points mark the tubercles forming a ‘V’. G, dorsal ornamentation of Karos tersum sp. nov. showing the transversal row of tubercles on mesotergal areas; transversal row II indicated by black dots. H, detail of the ventral ornamentation on femur IV of Karos barbarikos. I, detail of the ectal ornamentation on tibia IV of Karos barbarikos; arrows indicate the spiniform tubercles. J, detail of the ventral ornamentation on leg IV in Chapulobunus poblano sp. nov. K, detail of the two apical spines on femur IV in Chapulobunus poblano sp. nov. These images are not at the same scale.