Chersonesometrus nathanorum, Prendini & Loria, 2020

Prendini, Lorenzo & Loria, Stephanie F., 2020, Systematic Revision Of The Asian Forest Scorpions (Heterometrinae Simon, 1879), Revised Suprageneric Classification Of Scorpionidae Latreille, 1802, And Revalidation Of Rugodentidae Bastawade Et Al., 2005, Bulletin of the American Museum of Natural History 2020 (442), pp. 1-480 : 93

Chersonesometrus nathanorum , sp. nov.

Figures 10 , 16E , 30E , 61D , 62D , 63D , 80 , 99 ,

Figures

FIGURE 10. Phylogeny of Asian forest scorpions (Scorpionidae: Heterometrinae Simon, 1879, stat. nov.), obtained by simultaneous phylogenetic analysis of 186 morphological characters and 4188 aligned DNA nucleotides from three mitochondrial and two nuclear gene loci (Loria and Prendini, in press). Maximum likelihood tree with unambiguous morphological synapomorphies optimized. Black circles indicate uniquely derived apomorphic states, white circles parallel derivations of apomorphic states. Numbers above circles indicate characters, numbers below indicate states (appendix 1).

FIGURE 16. Chersonesometrus Couzijn, 1978, stat. nov. et stat. rev., carapace, dorsal aspect. A. Chersonesometrus bastawadei, sp. nov., holotype ♀ (AMNH), Naganath Temple, India. B. Chersonesometrus wroughtoni (Pocock, 1899), comb. nov., paralectotype ♀ (BMNH 1897.9.17.1-4, 11, 20). C, D. Chersonesometrus shivashankari, sp. nov., C. holotype ♂ (BMNH), Bellahunsi, India, D. paratype ♀ (BMNH), Hampasagar, India. E. Chersonesometrus nathanorum, sp. nov., subad. ♂ holotype (MNHN RS 3145). F. Chersonesometrus hendersoni, sp. nov., holotype ♀ (AMNH), Sanapura, India. Scale bars = 2 mm.

FIGURE 30. Chersonesometrus Couzijn, 1978, stat. nov. et stat. rev., sternum and pectines, ventral aspect. A. Chersonesometrus bastawadei, sp. nov., holotype ♀ (AMNH), Naganath Temple, India. B. Chersonesometrus wroughtoni (Pocock, 1899), comb. nov., paralectotype ♀ (BMNH 1897.9.17.1-4, 11, 20). C, D. Chersonesometrus shivashankari, sp. nov., C. holotype ♂ (BMNH), Bellahunsi, India, D. paratype ♀ (BMNH), Hampasagar, India. E. Chersonesometrus nathanorum, sp. nov., subad. ♂ holotype (MNHN RS 3145). F. Chersonesometrus hendersoni, sp. nov., holotype ♀ (AMNH), Sanapura, India. Scale bars = 2 mm.

FIGURE 61. Chersonesometrus Couzijn, 1978, stat. nov. et stat. rev., metasoma and telson, dorsal aspect. A. Chersonesometrus fulvipes (C.L. Koch, 1837), comb. nov., ♂ (BMNH 1896.7.30.91-92). B. Chersonesometrus shivashankari, sp. nov., paratype ♀ (BMNH), Hampasagar, India. C. Chersonesometrus madraspatensis (Pocock, 1900), comb. nov., ♂ (CAS 9071142). D. Chersonesometrus nathanorum, sp. nov., subad. ♂ holotype (MNHN RS 3145). E. Chersonesometrus wroughtoni (Pocock, 1899), comb. nov., paralectotype ♀ (BMNH 1897.9.17.1-4, 11, 20). Scale bars = 5 mm.

FIGURE 62. Chersonesometrus Couzijn, 1978, stat. nov. et stat. rev., metasoma and telson, lateral aspect. A. Chersonesometrus fulvipes (C.L. Koch, 1837), comb. nov., ♂ (BMNH 1896.7.30.91-92). B. Chersonesometrus shivashankari, sp. nov., paratype ♀ (BMNH), Hampasagar, India. C. Chersonesometrus madraspatensis (Pocock, 1900), comb. nov., ♂ (CAS 9071142). D. Chersonesometrus nathanorum, sp. nov., subad. ♂ holotype (MNHN RS 3145). E. Chersonesometrus wroughtoni (Pocock, 1899), comb. nov., paralectotype ♀ (BMNH 1897.9.17.1-4, 11, 20). Scale bars = 5 mm.

FIGURE 63. Chersonesometrus Couzijn, 1978, stat. nov. et stat. rev., metasoma and telson, ventral aspect. A. Chersonesometrus fulvipes (C.L. Koch, 1837), comb. nov., ♂ (BMNH 1896.7.30.91-92). B. Chersonesometrus shivashankari, sp. nov., paratype ♀ (BMNH), Hampasagar, India. C. Chersonesometrus madraspatensis (Pocock, 1900), comb. nov., ♂ (CAS 9071142). D. Chersonesometrus nathanorum, sp. nov., subad. ♂ holotype (MNHN RS 3145). E. Chersonesometrus wroughtoni (Pocock, 1899), comb. nov., paralectotype ♀ (BMNH 1897.9.17.1-4, 11, 20). Scale bars = 5 mm.

FIGURE 80. Map of South India and Sri Lanka, plotting known locality records of eight species of Chersonesometrus Couzijn, 1978, stat. nov. et stat. rev., based on material examined, and verified records from virtual museums and the literature: Chersonesometrus bastawadei, sp. nov.; Chersonesometrus beccaloniae (Kovařík, 2004), comb. nov.; Chersonesometrus hendersoni, sp. nov.; Chersonesometrus nathanorum, sp. nov.; Chersonesometrus pelekomanus (Couzijn, 1981), comb. nov.; Chersonesometrus shivashankari, sp. nov.; Chersonesometrus tristis (Henderson, 1919), comb. nov.; Chersonesometrus wroughtoni (Pocock, 1899), comb. nov.

FIGURE 99. Chersonesometrus nathanorum, sp. nov., subad. ♂ holotype (MNHN RS 3145), pedipalp patella: A. dorsal, B. retrolateral, and C. ventral aspects. Scale bar = 5 mm.