Cephennodes (Cephennodes) bilobatus, Jałoszyński, 2017

Jałoszyński, Paweł, 2017, Ant-like stone beetles on the roof of the world. Cephenniini of Nepal and Bhutan (Coleoptera, Staphylinidae, Scydmaeninae), Zootaxa 4349 (1), pp. 1-120 : 75-77

Cephennodes (Cephennodes) bilobatus sp. n.

( Figs 162 , 219–222 , 230 )

Type material. Holotype: NEPAL ( Sankhuwasabha District ): ♂, one label: "E. NEPAL: KOSI / Forêt S. Mangsingma / 2200 m, 11.IV.84 / Löbl - Smetana " [white, printed] ( MHNG) .

Diagnosis. BL 1.08 mm; abdominal sternite II with subtriangular, moderately broad and rounded lamina along posterior margin, sternite III with a similar but larger lamina; apex of median lobe of aedeagus subtriangular, blunt; apical projection of aedeagus distant from apex of median lobe, with distal margin shallowly concave in median region, strongly oblique in relation to the long axis of aedeagus, apical portion of apical projection broadly subtriangular.

Description. Body of male ( Fig. 162 ) brown, moderately stout, with shallow constriction between pronotum and elytra, strongly convex, covered with light brown setae; BL 1.08 mm.

Head subtrapezoidal, HL 0.15 mm, HW 0.30 mm; vertex and frons confluent and weakly, evenly convex, vertex with a pair of tiny but distinct tubercles; supraantennal tubercles small, weakly elevated; eyes moderately large, strongly convex, coarsely faceted. Punctures on vertex and frons small and shallow but distinct, moderately sharply marked, nearly evenly and densely distributed, separated by spaces slightly shorter than diameters of punctures; setae sparse, short, suberect. Antennae moderately long and relatively strongly thickened, AnL 0.58 mm, pentamerous club indistinctly delimited; antennomere I about 1.2 × as long as broad; II narrower than I but similar in length, 1.3 × as long as broad; III–VI each slightly narrower and much shorter than II, about as long as broad; VII slightly broader and indistinctly longer than VI, about as long as broad; VIII slightly broader than VII and similar in length, indistinctly transverse; IX and X each distinctly larger than preceding ones, each about as long as broad; XI distinctly broader than X, slightly shorter than IX–X together, about 1.5 × as long as broad.

Pronotum in dorsal view approximately semicircular, broadest near middle, PL 0.33 mm, PW 0.50 mm; anterior margin in strictly dorsal view nearly straight; lateral margins indistinctly microserrate, strongly rounded in anterior third, less so in posterior half; hind corners slightly obtuse-angled; posterior margin shallowly bisinuate; lateral carinae distinct but narrow, fused with lateral margins; antebasal pits small and shallow, each much closer to posterior than to lateral pronotal margin. Punctures on disc more distinct but similarly dense as those on frons and vertex, sharply marked; setae moderately long, dense, suberect.

Elytra elongate, oval, broadest slightly in front of middle, EL 0.60, EW 0.53, EI 1.14. Humeral calli weakly elevated; subhumeral lines distinct, divergent posteriorly, equal to about 0.4 EL, each developed as moderately sharp stepwise border between more convex humeral region and less convex adsutural area; basal pit on each elytron located in shallow and round basal impression; apices of elytra separately rounded. Punctures nearly as distinct and dense as those on pronotum; setae similar to those on pronotum.

Hind wings not studied.

Metaventrite with moderately large, sharply delimited postmesocoxal impressions, with median area covered with fine, dense punctures.

Abdomen ( Figs 219–220 ) strongly modified; sternite II with subtriangular, moderately broad and rounded lamina along posterior margin, sternite III with a similar but larger lamina.

Legs unmodified.

Aedeagus ( Figs 221–222 ) moderately slender, AeL 0.13 mm, apex of median lobe subtriangular; apical projection distant from apex of median lobe, its distal margin shallowly concave in median region, so strongly oblique in relation to the long axis of aedeagus that it becomes lateral margin; apex of apical projection broadly subtriangular; parameres very slender, subequal in length, not reaching apex of median lobe, each with one subapical seta.

Female. Unknown.

Distribution ( Fig. 230 ). Eastern Nepal.

Etymology. The name bilobatus refers to abdominal modifications, composed of two large subtriangular lobes.

Remarks. Within the C. simplicipes group, only C. bilobatus and C. bilaminatus have abdominal sternites II and III bearing laminae that strongly project posteroventrally and in lateral view visible as long subtriangular plates. Their shape, however, is clearly different; in C. bilobatus both laminae in ventral view are subtriangular and rounded, in C. bilaminatus subtrapezoidal and with straight posterior margins. Cephennodes bilobatus is also remarkably small; see remarks for a similarly small C. bahrabisensis .

Figures

FIGURES 156 – 162. Dorsal habitus of Cephennodes (s. str.) paralalitpuranus sp. n. (156), C. (s. str.) bahrabisensis sp. n. (157), C. (s. str.) bilaminatus sp. n. (158), C. (s. str.) ghorepanianus sp. n. (159), C. (s. str.) cordilaminatus sp. n. (160), C. (s. str.) mangmayanus sp. n. (161), and C. (s. str.) bilobatus sp. n. (162).

FIGURES 211 – 222. Abdomen of male in right lateral (211, 215, 219) and ventral (212, 216, 220) views and aedeagus in ventral (213, 217, 221) and lateral (214, 218, 222) views. Cephennodes (s. str.) cordilaminatus sp. n. (211 – 214), C. (s. str.) mangmayanus sp. n. (215 – 218), and C. (s. str.) bilobatus sp. n. (219 – 222).

FIGURES 229 – 230. Distribution of Cephennodes (s. str.) cordilaminatus sp. n. (229, circle), C. (s. str.) mangmayanus sp. n. (229, triangle), C. (s. str.) bilobatus sp. n. (230, circle).