Aleiodes (Hemigyroneuron) speciosus (Baker) Areekul-Butcher & Quicke, 2011

Areekul-Butcher, Buntika & Quicke, Donald L. J., 2011, Revision of Aleiodes (Hemigyroneuron) parasitic wasps (Hymenoptera: Braconidae: Rogadinae) with reappraisal of subgeneric limits, descriptions of new species and phylogenetic analysis, Journal of Natural History 45 (21 - 24), pp. 1403-1476 : 1458-1460

Aleiodes (Hemigyroneuron) speciosus (Baker) comb. nov.

( Figures 29 , 30 )

Hemigyroneuron speciosus Baker, 1917: 323 .

Material examined

Holotype. Female , Philippines, “Mt. Banahao, P.I., Baker; Hemigyroneuron speciosus Baker ; specimen figured C. van Achterberg, 1988 ” ( USNM).

Morphology

Length of body 8.5 mm, (remaining fore wing in two pieces).

Head. (Antenna incomplete); strongly transverse, 1.8 times wider than maximally long in dorsal view; width of head 3.0 times shortest distance between eyes; height of eye 1.67 times shortest distance between eyes; frons smooth and shiny, with some curved ridges paralleling the antennal sockets anteriorly, with short mid-longitudinal sulcus posteriorly, without pit; posterior ocellar line: transverse diameter of posterior ocellus: shortest distance between posterior ocellus and eye = 1.4: 5.0: 1.0; occiput finely rugulose superimposed on aciculate background; occipital carina complete.

Mesosoma . Mesoscutum coriaceous-rugulose and densely short-setose; notauli virtually undefined; mesopleuron with a large, slightly concave area of fine dorso-ventral striation, especially on its anterior half; prepectal carina incomplete, dorsally becoming confused in area of fine striation; scutellar sulcus with five carinae between the outer ones; scutellum with aciculate-coriaceous sculpture with disc margined by carina on anterior half becoming a faint irregular ridge posteriorly; propodeum coarsely foveate-reticulate superimposed on aciculate sculpture, with well-developed though rather irregular mid-longitudinal carina on anterior 0.7, coarsely crenulated along posterior margin, with slightly protruding postero-lateral tubercles but these not ridge-like.

Fore wing. Subbasal cell largely glabrous, without sclerome; posterior half swollen due to vein M+CU having an angulation at approximately 0.8 distance from base; vein 1-CU1, straight, 0.9 times length of 2-CU1; vein cu-a more or less vertical, straight, somewhat swollen anteriorly and narrowing posteriorly; subdiscal cell with reduced setosity antero-basally; length of veins r: 3-SR: SR1 = 1.0: 2.1: 5.0; vein 2-M approximately 2.5 times length of r-m; vein m-cu 2.0 times length of 2-SR+M, forming angle of 135 ◦ with 3-CU1.

Hind wing. Vein M+CU 1.4 times length of vein 1-M; vein 2-SC+ R tranverse; basal cell largely glabrous except posteriorly; subbasal and discal+subdiscal cells with large glabrous area anteriorly.

Claws. With three or four fine pectinal teeth on basal half of basal lobe.

Metasoma. Tergites 1, 2 and basal 0.25 of tergite 3 longitudinally striate with irregular punctures between the striae; 2nd tergite 1.42 times wider than medially long, 1.38 times longer than 3rd.

Colouration

Antennae black with flagellomeres 21– cream-white. Head largely ivory-white with clypeus, most of frons, vertex and occiput black. Mesosoma black with ivory-white marks on anterior and posterior of pronotum and propleuron, mid-posterior of mesoscutum extending as a “ V ”-shaped mark along notauli, large triangular posterodorsal mark on mesopleuron, most of metapleuron. Fore and mid coxa, trochanter and trochantellus, mid tarsus, hind trochanter, basal half of tibia and basal four tarsal segments ivory-white; hind coxa largely black and apex of hind femur black; rest of legs yellow-brown. Metasoma ivory-white with black transverse bands on middle of 1st tergite, near but removed from base of 2nd tergite, and broadly basally on 3rd– 6th tergites. Wings hyaline with dark brown venation except most of fore wing vein C+SC+ R and anterior margin of pterostigma brown-yellow (antennae incomplete).

Figures

Figure 29. Aleiodes (Hemigyroneuron) speciosus (Baker), CellD® light micrographs. (A) Lateral habitus; (B) head, dorsal aspect; (C) mesosoma, lateral aspect; (D) mesosoma, oblique dorsal aspect; (E) propodeum; (F) metasomal tergites 2–3.

Figure 30. Aleiodes (Hemigyroneuron) speciosus (Baker), CellD® light micrographs. (A) Remaining part of fore wing; (B) hind wing venation.