Alchemilla tzvelevii Czkalov, 2015

Chkalov, Andrey V., 2015, A new species of the genus Alchemilla (Rosaceae) from Central Russia, Phytotaxa 227 (3), pp. 282-288 : 284-285

Alchemilla tzvelevii Czkalov , sp. nov. (A. subsect. Alchemilla ser. Alchemilla subser. Heptagonae ). Fig. 1

Typus: — RUSSIA. Czuvaschia: distr. Kanasch, 2 km ad meridionali-orientem a pag. Malyje Bikschichi, pratula in declivi devexo expositione septentrionali-orientali depressionis, 55˚28ʹ19.9ʺ N, 047˚28ʹ57.6ʺ E, 13 June 2011, A. Czkalov & M. Makarov (holotypus [26/1]: LE!, isotypi: LE!, MW!, NNSU!). Fig. 2 .

Paratypi: — RUSSIA. Czuvaschia: distr. Kanasch, 2.5 km ad meridiem a pag. Malyje Bikschichi , pratulum (herbetum mesohygrophilum) in tilieto, 55˚27ʹ35.0ʺ N, 047˚29ʹ54.2ʺ E, 13 June 2011, A. Czkalov & M. Makarov ( LE!) ; ibidem: distr. Schemurscha, in vicinitas pag. Kuczeki , pratulum, 24 June 2010, M.M. Gafurova ( PVB!). Prov. Penza: distr. Kuznetsk, Aug 2001, M.S. Kulikovskij 037002 ; ibidem: 3 km ad orientem ab opp. Penza, prato humido, 06 May 1977, N. Jessina & M. Solovjova 037091 ( PKM!). Prov. Riazan’: distr. Spassk, 0.5 km ad meridiem a pag. Suschki, pratum secundum rivulum, 05 June 1994, M.V. Kazakova ( MW!) .

Planta flavo-viridis, paulo glaucescens, parva vel mediocris, caulibus ad 28 cm alt. Rhizoma sat crassum (1 cm cr.). Caules numero 2–4, erecti, foliis radicaliis aequales vel ea vix superantes, pilis paulo deflexis vel horizontaliter patentibus pubescentes, in dimidio superiore glabri vel pilis solitariis, interdum solum internodia infima pubescentes. Folia radicalia infima orbiculato-reniformia, sino angusto, angulo sectoris centrali ad 45°, fere plana, supra paulo aequaliter pubescentia, subtus nervis principalibus, dentibus, lobis extremis pubescentia, ceterum glabra; 5-lobata, lobis a 20%, zona centrali a 65% longitudinis folii (lg. f.) attengentibus, lobis planis vel semirotundatis obtusatis, incisura inter primum et centralem lobos ad 25% lg. f.; dentibus loborum utrinque 5 obtusiusculis, ovatis, fere aequilateralibus, obliquis, curvis, fere aequimagnis (lg. dentis unificati ad 10.5% lg. f., ad 53% lg. lobi), lg. dentis apicalis ad 75% lg. dentis unificati. Dentes laminae numero in toto a 70. Petioli foliorum tota longitudine glabri vel in dimidio superiore glabrescentes. Petioli foliorum radicalium ad 25 cm longi, laminae ad 5.5 cm lg., orbiculato-reniformes, orbiculate sino angusto vel lobis extremis tangentibus, angulo sectoris centrali 36–47°, planae vel paulo undulatae, laminae foliorum infimorum similiter pubescentes, sed nervis principalibus basi glabris, 7-lobata, lobis 24–26%, zona centrali 60.0–70.5 (–72.5)%, nervo primo (90–) 95–97%, nervo secundo (86–) 88–90% lg. f. attengentibus ; lobis trapezioideis, trapezioideo-semiovatis, lg. 43.0–51.0 (–54.5)%, distantia inter bases incisurarum 85–94 (–109)% lt. eorum; incisura inter primum et centralem lobos (6.2–) 8.0–11.0 (–13.5)% lg. f., 24.5–26.5% lg. lobi; dentibus loborum utrinque 7 obtusiusculis, aequilateralibus, ovatis, triangulari-ovatis, fere rectis, paulo curvis, fere aequimagnis (lg. dentis unificati 6.5–8.5% lg. f., 25.5–33.5% lg. lobi), lg. dentis apicalis (43–) 56–65% lg. dentis unificati). Lobi extremi 58–61 (– 67)% lg. f., (3–) 4–5 dentibus. Dentes laminae numero in toto 70–105. Petioli foliorum tota longitudine pilis paulo deflexis pubescentes, sub lamina saepe glabri. Stipulae foliorum radicalium purpurascentes, in sicco brunnescentes. Folium caulinum prope inflorescentiam singularem infimam late reniforme, petiolo 30–40%, zona centrali 45–60% lg. eius, 3-lobatum, dentibus loborum utrinque 2–3, lobis planis vel breviter trapezioideis, dentibus numero in toto 30–40. Synflorescentia angusta parviflora, 3–5 inflorescentiis singularibus. Inflorescentia singularis infima 0.5–1.0 lg. internodiae eius, ramulis glabris, floribus 10–20. Monochasia sat compacta. Monochasia laterales stipitibus reductis, axibus brevibus (ad 0.5 lg. pedicelli), floribus 3–4. Monochasia terminale stipitibus sat brevibus (circa 1 lg. pedicelli), axibus aperte (1–2 lg. pedicelli), floribus 5–6. Flores mediocres, hypanthia obconica, rarius tubulosa. Hypanthia matura campanulata, saepe orciformia vel rotundata, cum collo, glabra, (1.27–) 1.46–1.83 (–2.07) mm lg., eorum relatio latitudo (lt.): lg. (0.69–) 0.71–0.78 (–0.85). Pedicelli 1–2 lg. hypanthii, glabri, incrassatione cacuminis aperta. Sepala triangulari-ovata, (1.19–) 1.27–1.51 (–1.59) mm lg., (67–) 76–83 (–100)% lg. hypanthii, eorum relatio lt.: lg. (0.46–) 0.64–0.76, glabra vel barbellata. Episepala lanceolata, (0.73–) 0.83–1.15 (–1.27) mm lg., (47–) 55–64 (–69)% lg. hypanthii, (58–) 68–78 (–84)% lg. sepalorum, eorum lt. (38–) 42–51% lt. sepalorum, eorum relatio lt.: lg. 0.35–0.44 (–0.67), glabra vel pilis solitariis.

Habitat in pratis subhumidis et inundatis, marginibus silvarum frondosarum in regionibus partis mediae fluminis Volga.

Affinitas. Species hic characteres grecum valde distantium complectitur. Quae habituale, dissectioni folii, incisurae profundae inter lobos, dentibus magnis, numero pauco dentium folii, monochasiis sat compactis, formae hypanthii speciebus A. subsect. Pubescentes, quidem A. hirsuticauli , valde similis est. Toti his characteri species nostra ab speciebus e A. subser. Heptagonae ( A. devestiens Juzepczuk (1951: 165) , A. decalvans Juzepczuk (1933: 125) etc.), ad quam formale referatur, optime differt.

Figures

FIGURE 1. Main morphological characteristics of Alchemilla tzvelevii. A, B. Lamina of the upper rosette leaf. C. Lamina of the middle rosette leaf. D. Terminal monochasium. E. Lateral monochasium. F. Indumentum of petiole. G. Hypanthia in bloom. H. Hypanthia in fruits. Scale bar = 5 mm.

FIGURE 2. Alchemilla tzvelevii, holotype. Scale bar = 10 cm.