Xiphocentron (Sphagocentron) tapanti, Vilarino & Calor, 2024

Vilarino, Albane & Calor, Adolfo R., 2024, Comparative geometric morphometrics of male genitalia in Xiphocentron subgenera (Trichoptera: Xiphocentronidae): new species, revision and phylogenetic systematics of the subgenus Sphagocentron, Arthropod Systematics & amp; Phylogeny 82, pp. 407-431 : 407-431

publication ID

https://doi.org/ 10.3897/asp.82.e112587

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Arthropod Systematics & Phylogeny by Pensoft

scientific name

Xiphocentron (Sphagocentron) tapanti

sp. nov.

Xiphocentron (Sphagocentron) tapanti sp. nov.

Figures 11 A – D View Figure 11

Type material.

Holotype COSTA RICA • ♂; Cartago; Tapanti Reserve , quebrada palmitos and falls, 9.72 ° N, 83.78 ° W, 24–25. iii. 1991, el. 1400 m, Holzenthal, Muñoz, Huisman leg., UMSP 000143451 View Materials GoogleMaps . — Paratypes COSTA RICA • 2 ♂; same data as holotype, UMSP 000143450 View Materials , 000143452 View Materials GoogleMaps . • 9 ♂; same data, except 23. viii. 1990, Holzenthal and Huisman leg., UMSP 000143453 View Materials , 000143454 View Materials , 000143455 View Materials , 000143456 View Materials , 000143457 View Materials , 000143458 View Materials , 000143462 View Materials , 000143463 View Materials , 000143464 View Materials GoogleMaps .


The new species is similar to Xiphocentron julus by the posterior margin of sternum IX with weakly produced mesal lobes. It can be differentiated from these and other species by the combination of the following characters: (1) the preanal appendage very wide subapically (narrow throughout length in X. julus ); (2) shape of paraproct in lateral view, with apicodorsal margin truncate with conspicuously narrow ventral lobe (apicodorsal margin oblique, contiguous with the ventral lobe in X. julus ); (3) tergun IX in dorsal view, posterior and anterior margins with wide deep mesal incisions.


Male: Forewing length 3.6–4.4 mm (n = 12). Color overall brown, forewing uniformly dark brown. Maxillary palp segment length formula (I = II = III) <IV <V. Tibial spur formula 2: 4: 3; spurs unmodified. Forewing forks II and IV present; fork II sessile at discoidal cell; discoidal 3 / 4 as long as thyridial cell. Hind wing forks II and V present (Fig. 11 A View Figure 11 ). Sternum V with short, flattened projection, somewhat angular apically.

Genitalia (Fig. 11 B – E View Figure 11 ). Tergum IX, in lateral view, wide basally, narrower apically (Fig. 11 B View Figure 11 ); in dorsal view, anterior margin with deep concave mesal incision; posterior margin with round lobes and deep mesal incision (Fig. 11 C View Figure 11 ). Sternum IX, in lateral view, about 3 × as long as high, apex subround; anterior apodeme wide, straight, slightly tapering (Fig. 11 B View Figure 11 ); in ventral view, slightly longer than wide, posterior margin substraight, with shallow, narrow mesal incision forming weakly produced mesal lobes (Fig. 11 D View Figure 11 ). Tergum X membranous fused basodorsally to each paraproct. Paraproct, in lateral view, oblong, wide at base, narrowing apically, with narrow, ventral lobe (Fig. 11 B View Figure 11 ); in dorsal view, wide basally, divided apicomesally, each side fused at midlenght, without sclerotized mesal band, apex with several sensillae (Fig. 11 C View Figure 11 ). Preanal appendage setose, in lateral view about 2.5 × as long as tergum IX, sigmoid, narrow at base, wide subapically, about 2 × as wide as base, tapering at apex (Fig. 11 B View Figure 11 ); in dorsal view, substraight, tapering apically (Fig. 11 C View Figure 11 ). Inferior appendage, in lateral view, about 2.5 × as long as tergum IX; coxopodite and harpago fused, with suture line between each article, inner face basally with long, dense spine-like setae, and row of regular setae from midlenght to apex (Fig. 11 B View Figure 11 ); basal region (coxopodite) wide, without basomesal spine-like setae, mesal sclerite region with few difuse long spine-like setae; apical region (harpago) narrow, digitate, about as long as basal region, apex slightly enlarged, round (Fig. 11 B View Figure 11 ); basal plate in lateral view wide, anteriorly with, narrow flange (Fig. 11 B View Figure 11 ). Phallus tubular, very long and narrow, reaching segment V; basally conical, subapically annulate, weakly sclerotized; apex slightly enlarged (Fig. 11 E View Figure 11 ).


Name in apposition; in reference to the Tapantí National Park where the species was collected.


Costa Rica.