Ettingshausenia? lublinensis ( Karczmarz and Popiel, 1971 ) Halamski, 2013

Halamski, Adam T., 2013, Latest Cretaceous leaf floras from southern Poland and western Ukraine, Acta Palaeontologica Polonica 58 (2), pp. 407-443 : 425

Ettingshausenia? lublinensis ( Karczmarz and Popiel, 1971) comb. nov.

Figs. 11C, E.

1907 Platanus affinis (?) Lesqx.; Nowak 1907a: 51, pl. 1: 1.

1971 Platanus lublinensis ; Karczmarz and Popiel 1971: 647–648, pl. 3: 9, 4: 10,?11.

Basionym : Platanus lublinensis Karczmarz and Popiel, 1971 .

Type material: Holotype JS1, large part of a leaf figured by Karczmarz and Popiel (1971: pl. 3: 9) and re−figured herein ( Fig. 7C ); paratype, a fragmentary leaf figured by Karczmarz and Popiel (1971: pl. 4: 10) (collection number 163, specimen not traced). The second paratype figured by Karczmarz and Popiel (1971: pl. 4: 11) is a loose bark impression and therefore unrecognisable.

Material.— Holotype and a single fragmentary leaf L PB−K.2 from Potelych.

Description.—Leaf and blade attachment not preserved. Laminar size mesophyll (length of the preserved part 23 cm, estimated total length ca. 30 cm), laminar length to width ratio about 1.25:1, laminar shape obovate, shallowly palmately trilobed (the unlobed part represents about 80% of the total leaf length); margin slightly undulating but mostly entire. Apex of the lateral lobe flattened, that of the median lobe not preserved. Base not preserved.

Primary veins three, basal actinodromous. Secondary veins craspedodromous, their spacing decreasing distally, departing at about 20 ° (thus parallel to leaf margins); the angle of departure is rather constant throughout the entire leaf, while the secondaries bend inwards in the distal parts of the leaf. Interior secondaries probably absent. Intercostal tertiary vein fabric alternate percurrent; epimedial tertiaries percurrent; exterior tertiaries looped. Quaternary vein fabric irregularly reticulate; quinquenary vein fabric poorly preserved, probably similar to the precedent.

Remarks.—The discussed species is tentatively placed within the form genus Ettingshausenia Stiehler, 1857 on account of pronounced lobes and probable absence of suprabasal veins ( Kvaček and Váchová 2006). As a matter of fact, the base of the leaf is not preserved but the relatively narrow surface of the blade between the leaf margin and the lateral primary vein is rather suggestive of a leaf organisation without suprabasal veins. Cenomanian Ettingshausenia bohemica (Velenovský, 1882) Kvaček and Váchová, 2006 is the nearest species, differing, however, from Ettingshausenia? lublinensis in much wider blade, more marked serration, and more pronounced lobes ( Knobloch 1997; Kvaček and Váchová 2006). Ettingshausenia senoniana ( Knobloch, 1964) Kvaček and Váchová 2006 from the Senonian of Bohemia and E. gruenbachiana Herman and Kvaček, 2010 from the lower Campanian of Grünbach in Austria ( Herman and Kvaček 2010) both have more acute lobes.

Figures

Fig. 7. A, C. Eudicot Debeya insignis (Hosius and von der Marck, 1880) Knobloch, 1964. A. Lectotype PBO 1021[−1]; Haldem, Stemweder Berg, upper Campanian. General view (A1), enlargement of the petiolule branching (A2). C. General view of the slab PBO 1021 showing the lectotype (shown in A) and a paralectotype (PBO 1021[−2]) of Debeya insignis as well as two leaflets of Debeya haldemiana (shown in B); Haldem, Stemweder Berg, upper Campanian. B, G. Debeya haldemiana de Saporta and Marion, 1873. B. Fragmentary leaf PBO 1021[−3]; Haldem, Stemweder Berg, upper Campanian. G. Isolated leaflet ZPAL 4/2/2; Bliżów, lower Maastrichtian. D. Eolirion sp., fragmentary leaf ZPAL Pl 4/1/9; Krasbobród, upper Campanian. E. Debeya paulinae sp. nov., fragmentary leaf KrM Pb 1/6 (paratype); Krasnobród, upper Campanian. F. Debeya paulinae sp. nov., paratype KrS 1, enlargement of the petiolule branching (see general view in Fig. 2C). Scale bars 10 mm.