Zeanillus pallidus (Broun),

Sokolov, Igor M., 2016, A taxonomic review of the anilline genus Zeanillus Jeannel (Coleoptera: Carabidae: Bembidiini) of New Zealand, with descriptions of seven new species, re-classification of the species, and notes on their biogeography and evolution, Zootaxa 4196 (1), pp. 1-37: 16-17

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Zeanillus pallidus (Broun)


Zeanillus pallidus (Broun) 

Figs. 1View FIGURE 1 C, 2B, 3C –D, 4C, 5C, 6C, 7D, 8B, 10DIO, 11D, 12M –P, 14D, 15D, 16E, 18A

Anillus pallidus Broun, 1884: 228  . Type locality: «Sea Beach, Taieri» (original citation). Anillus marginatus Broun, 1914: 152  . Type locality: «Invercargill» (original citation). Synonymy established by Moore (1980: 402).

Examined material. More than one hundred specimens (15 dissected) from the following localities: Otago: Symes Road at Old Man Range, 10km S Beaumont, Flagstaff-Whare Flat Road near Dunedin, Taieri Mouth, Picnic Gulley, Akatore  ; Southland: Otanomomo Reserve, Kaka Point Scenic Reserve, Titpua near Marbel Bush, Thomsons Bush in Invercargill. 

Recognition. Adults of this species ( Fig. 11View FIGURE 11 D) are distinguished from other congeners by the characters listed in the subgeneric diagnosis.

Description. Size. Medium for genus (SBL range 1.62–1.84 mm, mean 1.73± 0.054 mm, n=20).

Habitus. Body form subdepressed, subparallel, moderately elongate (WE/SBL 0.34±0.006), head of moderate width for genus compared to pronotum (WH/WPm 0.77±0.018), pronotum of moderate width in comparison to elytra (WPm/WE 0.76±0.014).

Color. Body color testaceous, appendages testaceous.

Prothorax. Pronotum slightly long (LP/LE 0.39±0.012) and moderately elongate (WPm/LP 1.29±0.028), with lateral margins rectilinear and markedly constricted posteriorly (WPm/WPp 1.97±0.076). Width between anterior angles much greater than between posterior angles (WPa/WPp 1.51±0.069).

Elytra. Moderately depressed, comparatively long (LE/SBL 0.51±0.011) and slightly wide (WE/LE 0.67±0.015). Lateral margins slightly divergent at basal fourth, subparallel at middle and evenly rounded to apex in apical third.

Male genitalia. Median lobe ( Fig. 12View FIGURE 12 M) with straight ventral margin and semicircular short apex. Apical orifice area without ostial fields. Sclerites of internal sac united basally, of moderate length, much shorter than the length between apical and basal orifices. Left paramere without apical constriction ( Fig. 12View FIGURE 12 O). Ring sclerite with short and markedly transverse handle ( Fig. 14View FIGURE 14 D).

Female internal genitalia. Goncoxite 2 with short ensiform setae ( Fig. 15View FIGURE 15 D). Spermatheca sclerotized, elongate, with two compartments ( Fig. 16View FIGURE 16 E). Spermathecal duct short and without coils. Attachments of spermathecal duct and gland to spermatheca close together.

Geographical distribution. This species inhabits coastal plains of the eastern Southland and South Otago, from Invercargill up to Dunedin, also goes upstream along the Clutha River almost to Alexandra in Central Otago (18A, green diamonds).

Way of life. Specimens were collected in broadleaf/podocarp and Nothofagus  forests.

Relationships. The structure of male genitalia suggest the relatedness to the representatives of Zeanillus  s.str. The absence of postorbital seta is a character shared with the members of Otagonillus. Based on these facts and distributional data it is postulated that Z. pallidus  is the sister taxon to the members of Zeanillus  s.str. and shared a common ancestry with the representatives of Otagonillus.














Zeanillus pallidus (Broun)

Sokolov, Igor M. 2016

Anillus pallidus

Moore 1980: 402
Broun 1914: 152
Broun 1884: 228