Broun, 1884 : 228 Broun, 1914 : 152 Moore (1980: 402) A taxonomic review of the anilline genus Zeanillus Jeannel (Coleoptera: Carabidae: Bembidiini) of New Zealand, with descriptions of seven new species, re-classification of the species, and notes on their biogeography and evolution Sokolov, Igor M. Zootaxa 2016 4196 1 1 37 8MF5D Broun Broun [151,474,1122,1149] Insecta Carabidae Zeanillus Animalia Coleoptera 15 16 Arthropoda species pallidus     Anillus pallidus  Broun, 1884: 228. Type locality: «Sea Beach, Taieri» (original citation).  Anillus marginatus  Broun, 1914: 152. Type locality: «Invercargill» (original citation). Synonymy established by  Moore (1980: 402).    Examined material.More than one hundred specimens (15 dissected) from the following localities: Otago: Symes Road at Old Man Range,  10kmS Beaumont, Flagstaff-Whare Flat Roadnear Dunedin, Taieri Mouth, Picnic Gulley, Akatore;  Southland: Otanomomo Reserve, Kaka Point Scenic Reserve, Titpua near Marbel Bush, Thomsons Bush in Invercargill.  Recognition.Adults of this species ( Fig. 11D) are distinguished from other congeners by the characters listed in the subgeneric diagnosis.   Description.Size. Medium for genus (SBL range 1.62–1.84 mm, mean 1.73± 0.054 mm, n=20). Habitus. Body form subdepressed, subparallel, moderately elongate (WE/SBL 0.34±0.006), head of moderate width for genus compared to pronotum (WH/WPm 0.77±0.018), pronotum of moderate width in comparison to elytra (WPm/WE 0.76±0.014). Color. Body color testaceous, appendages testaceous. Prothorax. Pronotum slightly long (LP/LE 0.39±0.012) and moderately elongate (WPm/LP 1.29±0.028), with lateral margins rectilinear and markedly constricted posteriorly (WPm/WPp 1.97±0.076). Width between anterior angles much greater than between posterior angles (WPa/WPp 1.51±0.069). Elytra. Moderately depressed, comparatively long (LE/SBL 0.51±0.011) and slightly wide (WE/LE 0.67±0.015). Lateral margins slightly divergent at basal fourth, subparallel at middle and evenly rounded to apex in apical third. Male genitalia. Median lobe ( Fig. 12M) with straight ventral margin and semicircular short apex. Apical orifice area without ostial fields. Sclerites of internal sac united basally, of moderate length, much shorter than the length between apical and basal orifices. Left paramere without apical constriction ( Fig. 12O). Ring sclerite with short and markedly transverse handle ( Fig. 14D). Female internal genitalia. Goncoxite 2 with short ensiform setae ( Fig. 15D). Spermatheca sclerotized, elongate, with two compartments ( Fig. 16E). Spermathecal duct short and without coils. Attachments of spermathecal duct and gland to spermatheca close together.  Geographical distribution.This species inhabits coastal plains of the eastern Southlandand South Otago, from Invercargill up to Dunedin, also goes upstream along the Clutha River almost to Alexandra in Central Otago(18A, green diamonds).  Way of life.Specimens were collected in broadleaf/podocarp and  Nothofagusforests.  Relationships.The structure of male genitalia suggest the relatedness to the representatives of  Zeanillus s.str.The absence of postorbital seta is a character shared with the members of Otagonillus. Based on these facts and distributional data it is postulated that  Z. pallidusis the sister taxon to the members of  Zeanillus s.str.and shared a common ancestry with the representatives of Otagonillus. 1341350632 Flagstaff-Whare Flat Road 15 16 1 Otago 1341350617 Southland 15 16 1 Southland