Tafana humahuaca, De Oliveira & Brescovit, 2021

De Oliveira, Luiz Fernando M. & Brescovit, Antonio Domingos, 2021, Taxonomic revision and cladistic analysis of ghost spiders of the genus Tafana Simon, 1903 (Araneae: Dionycha, Anyphaenidae), with the descriptions of twelve new species, European Journal of Taxonomy 742, pp. 1-77 : 25-27

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scientific name

Tafana humahuaca

sp. nov.

Tafana humahuaca sp. nov.


Figs 15 View Fig , 32J View Fig , 35 View Fig


Males of Tafana humahuaca sp. nov. resemble those of T. riveti and T. nevada sp. nov. by having the short embolus, but differ by having the apophysis at the base of the embolus truncated, extremely short and with excavation between the base of the embolus and the apophysis ( Fig. 15C–E View Fig ). Females resemble those of Tafana nevada sp. nov. by having the epigynal plate, but differ by having atrium narrow in the anterior region, small hood with lateral lobes extremely curved in the anterior region ( Figs 15F–G View Fig , 32J View Fig ).


The specific name, a noun in apposition, is a tribute to ‘Quebrada de Humahuaca’, a deep canyon located in Jujuy Province, the type locality.

Material examined

Holotype ARGENTINA • 1 ♂; Jujuy Province, Santa Bárbara; [24°18′31″ S, 64°39′37″ W]; Oct. 1969; G. Williner leg.; MACN-Ar 41787 . GoogleMaps

Paratype ARGENTINA • 1 ♀; same collection data as for holotype; Oct. 1962; MACN-Ar 41788 GoogleMaps .


Male (holotype, MACN-Ar 41787)

COLOR. Exemplar probably discolored. Carapace orange ( Fig. 15A View Fig ). Labium, sternum and endites yellow. Chelicerae yellow. Legs I–III yellow with spots on the articles. Abdomen with dark spots ( Fig. 15A View Fig ).

MEASUREMENTS. Total length 5.3, carapace length 2.3, width 1.7. Clypeus height 0.06. Eye diameters and interdistances: AME 0.06, ALE 0.1, PME 0.1, PLE 0.08; AME–AME 0.02, AME–ALE 0.02, PME– PME 0.1, PME–PLE 0.08, ALE–PLE 0.02. Chelicerae 0.8 long, with five promarginal teeth and four retromarginal denticles.

LEG MEASUREMENTS. I—lost; II—femur 1.7/ patella 0.73 /tibia 1.7/ metatarsus 1.6 /tarsus 0.84 / total 6.57; III—1.6 / 0.7/ 1.4 /1.4 / 0.7/ 5.8; IV—lost.

LEG SPINATION. II—tibia v2-1-2, p1-1-0; III—tibia v1-2-0, p0-1-0, metatarsus v2-1-0, p0-1-0.

PALP. Retrolateral tibial apophysis conical with apex curved; cymbium with retrolateral projection more rounded; embolus short; ventral tegular process near the median apophysis and sclerotized; sperm duct in vertical position with secondary looping reaching the apex of the tegulum; median apophysis slender and curved apically ( Fig. 15C–E View Fig ).

ABDOMEN. Length 2.7, epigastric furrow 0.6 from tracheal spiracle, spiracle 1.0 from base of spinnerets.

Female (paratype, MACN-Ar 41788)

COLOR. Exemplar probably discolored. Coloration as in males of this species ( Fig. 15B View Fig ).

MEASUREMENTS. Total length 3.2, carapace length 2.4, width 1.3. Clypeus height 0.06. Eye diameters and interdistances: AME 0.08, ALE 0.1, PME 0.1, PLE 0.1; AME–AME 0.04, AME–ALE 0.04, PME– PME 0.16, PME–PLE 0.12, ALE–PLE 0.04. Chelicerae 1.1 long, with four promarginal teeth and five retromarginal denticles.

LEG MEASUREMENTS. I—femur 1.6 /patella 0.8/ tibia 1.4/ metatarsus 1.3/ tarsus 0.8 /total 5.9; II— 1.9/ 0.8/ 1.1 /1.15 / 0.6 /5.55; III—1.3/ 0.63 / 0.8/ 1.2/ 0.5/ 4.43; IV—1.9 / 0.8/ 1.5/ 2.1/ 0.65 / 6.95.

LEG SPINATION. I—tibia v2-2-0, metatarsus v2-0-0; II—tibia v1-1-0; III—tibia v1-2-2, p1-1-0, r1-1-0, metatarsus v2-1-2; IV—tibia v1-2-2, p1-1-1.

EPIGYNUM. Hood semicircular and sclerotized; short atrium; lateral lobes over copulatory ducts. Internally, copulatory ducts narrow; primary spermathecae large and oval; secondary spermathecae above the primary spermathecae and rounded; fertilization ducts long ( Fig. 15F–G View Fig ).

ABDOMEN. Length 3.2, epigastric furrow 0.7 from tracheal spiracle, spiracle 1.3 from base of spinnerets.


Known only from Jujuy, Argentina ( Fig. 35 View Fig ).