Taxonomic revision and cladistic analysis of ghost spiders of the genus Tafana Simon, 1903 (Araneae: Dionycha, Anyphaenidae), with the descriptions of twelve new species Author De Oliveira, Luiz Fernando M. 06C42876-2925-4007-A19D-053BD0058F25 Laboratório de Coleções Zoológicas, Instituto Butantan - Av. Vital Brazil, 1500, 05503 - 900, São Paulo, SP, Brazil. & Instituto de Biociências, Universidade de São Paulo, Rua do Matão, 321, 05508 - 090, São Paulo, SP, Brazil. luiz.moura.lfm@gmail.com Author Brescovit, Antonio Domingos D5B81D79-AFAE-47B1-8A6E-DAB448A24BCC Laboratório de Coleções Zoológicas, Instituto Butantan - Av. Vital Brazil, 1500, 05503 - 900, São Paulo, SP, Brazil. antonio.brescovit@butantan.gov.br text European Journal of Taxonomy 2021 2021-03-31 742 1 77 http://dx.doi.org/10.5852/ejt.2021.742.1291 journal article 10.5852/ejt.2021.742.1291 62f9dc57-bade-4fbc-8af7-f92ff107593c 2118-9773 4655404 23424EC3-8440-4C72-AD9E-C1EC3087F7CA Tafana humahuaca sp. nov. urn:lsid:zoobank.org:act: 16D23428-F7B9-48A6-93EE-95B3AE8F97DC Figs 15 , 32J , 35 Diagnosis Males of Tafana humahuaca sp. nov. resemble those of T. riveti and T. nevada sp. nov. by having the short embolus, but differ by having the apophysis at the base of the embolus truncated, extremely short and with excavation between the base of the embolus and the apophysis ( Fig. 15C–E ). Females resemble those of Tafana nevada sp. nov. by having the epigynal plate, but differ by having atrium narrow in the anterior region, small hood with lateral lobes extremely curved in the anterior region ( Figs 15F–G , 32J ). Etymology The specific name, a noun in apposition, is a tribute to ‘Quebrada de Humahuaca’, a deep canyon located in Jujuy Province , the type locality. Material examined Holotype ARGENTINA1 ♂ ; Jujuy Province , Santa Bárbara ; [ 24°18′31″ S , 64°39′37″ W ]; Oct. 1969 ; G. Williner leg.; MACN-Ar 41787 . Paratype ARGENTINA1 ♀ ; same collection data as for holotype; Oct. 1962 ; MACN-Ar 41788 . Description Male ( holotype , MACN-Ar 41787) COLOR. Exemplar probably discolored. Carapace orange ( Fig. 15A ). Labium, sternum and endites yellow. Chelicerae yellow. Legs I–III yellow with spots on the articles. Abdomen with dark spots ( Fig. 15A ). MEASUREMENTS. Total length 5.3, carapace length 2.3, width 1.7. Clypeus height 0.06. Eye diameters and interdistances: AME 0.06, ALE 0.1, PME 0.1, PLE 0.08; AME–AME 0.02, AME–ALE 0.02, PME– PME 0.1, PME–PLE 0.08, ALE–PLE 0.02. Chelicerae 0.8 long, with five promarginal teeth and four retromarginal denticles. LEG MEASUREMENTS. I—lost; II—femur 1.7/ patella 0.73 /tibia 1.7/ metatarsus 1.6 /tarsus 0.84 / total 6.57; III—1.6 / 0.7/ 1.4 /1.4 / 0.7/ 5.8; IV—lost. LEG SPINATION. II—tibia v2-1-2, p1-1-0; III—tibia v1-2-0, p0-1-0, metatarsus v2-1-0, p0-1-0. PALP. Retrolateral tibial apophysis conical with apex curved; cymbium with retrolateral projection more rounded; embolus short; ventral tegular process near the median apophysis and sclerotized; sperm duct in vertical position with secondary looping reaching the apex of the tegulum; median apophysis slender and curved apically ( Fig. 15C–E ). ABDOMEN. Length 2.7, epigastric furrow 0.6 from tracheal spiracle, spiracle 1.0 from base of spinnerets. Female ( paratype , MACN-Ar 41788) COLOR. Exemplar probably discolored. Coloration as in males of this species ( Fig. 15B ). MEASUREMENTS. Total length 3.2, carapace length 2.4, width 1.3. Clypeus height 0.06. Eye diameters and interdistances: AME 0.08, ALE 0.1, PME 0.1, PLE 0.1; AME–AME 0.04, AME–ALE 0.04, PME– PME 0.16, PME–PLE 0.12, ALE–PLE 0.04. Chelicerae 1.1 long, with four promarginal teeth and five retromarginal denticles. LEG MEASUREMENTS. I—femur 1.6 /patella 0.8/ tibia 1.4/ metatarsus 1.3/ tarsus 0.8 /total 5.9; II— 1.9/ 0.8/ 1.1 /1.15 / 0.6 /5.55; III—1.3/ 0.63 / 0.8/ 1.2/ 0.5/ 4.43; IV—1.9 / 0.8/ 1.5/ 2.1/ 0.65 / 6.95. LEG SPINATION. I—tibia v2-2-0, metatarsus v2-0-0; II—tibia v1-1-0; III—tibia v1-2-2, p1-1-0, r1-1-0, metatarsus v2-1-2; IV—tibia v1-2-2, p1-1-1. EPIGYNUM. Hood semicircular and sclerotized; short atrium; lateral lobes over copulatory ducts. Internally, copulatory ducts narrow; primary spermathecae large and oval; secondary spermathecae above the primary spermathecae and rounded; fertilization ducts long ( Fig. 15F–G ). ABDOMEN. Length 3.2, epigastric furrow 0.7 from tracheal spiracle, spiracle 1.3 from base of spinnerets. Distribution Known only from Jujuy , Argentina ( Fig. 35 ).