Procerini

Nygren, Arne, 2004, Revision of Autolytinae (Syllidae: Polychaeta)., Zootaxa 680, pp. 1-314 : 31-39

Nuchal epaulettes extending to half of chaetiger 2, large eye spots .................................. ................................................................................................... M. rangiroaensis (p. 155) Nuchal epaulettes extending to between end of chaetiger 3 and beginning of chaetiger 5, small eye spots ................................................................................... M. dentalia (p. 127) 85 (83) Trepan with 4–5 large and 25–39 small teeth, 1 large alternating with 4–13 small. Small teeth fused with adjacent large teeth ........................................... M. langerhansi (p. 140) Trepan with 8–10 large and 14–21 small teeth, 1 large alternating with 1–3 small. Small teeth not fused with adjacent large teeth ................................ M. brachycephala (p. 120) Procerini trib. n.

Pharynx sinuation Trepan teeth Proventricle in chaetiger Reproduc­ tive mode Additional fea­ tures Distri­ bution References branchiata 1– several 1 20 uneq; 1L+(1–3)S 6–8(2–3) 50 S; AS Furcating nuchal epaulettes ANT 1Averincev, 1972 clavata several 24 eq 5(1) 20 S; AS NEA longi­ pharyngea several 20 eq 4–5(1) 18? SWP pectinans 1 217 uneq; 1L+(1–2)S 5–6(2) 26 S; AS EIO 2Hartmann­ Schröder, 1989 spirifer 1 28 uneq; 1L+(1–3)S 3(1) 25 S; AS SWP,

Rows of muscle cells –30 EIO

Linnean name definition. Type genus Proceraea Ehlers, 1864 .

Node­based name definition. Procerini refers to the least inclusive clade comprising Imajimaea zonata ( Mohammad, 1973) , Imajimaea draculai ( San Martín & López, 2002), Virchowia [ Langerhans, 1879] Nygren, 200X, Levidorum [ Hartman, 1967] Nygren, 200X, and Proceraea [ Ehlers, 1864] Nygren, 200X.

Apomorphies. Clade supported by six morphological apomorphies ( Fig. 4 ): 1) nuchal epaulettes extend to end of tentacular segment (character 8), character state change is a parallelism, character later reversed within clade, character further changed within clade, character state unknown in some taxa; 2) cirrostyles alternate in length present (character 16), character state change is a parallelism, character later reversed within clade, character state unknown in some taxa; 3) cirrophores on short cirri absent (character 17), character state change is a parallelism, character later reversed within clade, character state unknown in some taxa; 4) cirrophores on long cirri absent (character 18), character state change is a parallelism, character later reversed within clade, character state unknown in some taxa; 5) chaetal teeth in median and posterior chaetigers with distal tooth smaller than subdistal tooth (character 31), character state change is a parallelism, character later reversed within clade, character state unknown in some taxa; 6) reproductive mode equals anterior scissiparity (character 51), other MPRs possible, character state unknown in some taxa.

Figures

FIGURE 4. Character transformations. Symbols as follow: black box, only one most parsimonious reconstruction (MPR) exists; white box, more than one MPR exist; X reversal; = parallelism; character later reversed within clade; character further changed within clade;? unknown character state for one or several taxa within clade.

Tables

TABLE 8 A. A selection of characters for Myrianida ­ taxa. Colour refers to live specimens. Abbreviations as occuring under each headline. Length of the proventricle is noted in brackets and measured as the number of body segments it covers. Nuchal epaulettes: b = beginning, m = middle, e = end, ts = tentacular segment, 1 = chaetiger 1. Ets equals extension of nuchal epaulettes to the end of tentacular segment. Trepan teeth, teeth arrangement: L = large, S = small. Basal ring, infradental spines: y = present, n = absent.

	Colour	Nuchal epua­ lettes	Pharynx sinua­ tion	Trepan in chaetiger	Trepan teeth	Teeth arrange­ ment	Basal ring	Infradental spines
arborea	uncoloured 1 m1	m1	1	3–4	c. 18; eq		y	y
australiensis	?	e3	1, sev­ eral	1	16–18; eq		thin	y
brachycephala	uncoloured to faintly orange	e1–e2	1	2–4	22–29; (8– 10)L+(14–21)S	1L+(1–3)S	y	y
brevicirrata	?	e1	1	2	26–30; eq		thin	y
brevipes	?	ets	1	1	20–25; eq		y?	y?
convoluta	uncoloured	e1–e2	several	1–2	9, 15–16; eq		thin	y
dentalia	uncoloured to faintly pinkish anteriorly	e3–b5	1	3–7	27–40; 2L+(25– 38)S	1L+(9–11 dor­ sal)+1L+ (16–19 ventral)	y	y
edwarsi	reddish sides anteriorly	b1–b2	1	1–3	24–34; eq		y	y

...... continued on the next page