Bengalia peuhi Villeneuve, 1914

Rognes, Knut, 2012, 3553, Zootaxa 3553, pp. 1-79 : 14-20

2. Bengalia peuhi Villeneuve, 1914

Figs. 20–42.

Auchmeromyia peuhi Brauer & Bergenstamm, 1891: 420 [as “ Peuhi Mcq. Senegal ” under “ Auchmeromyia S. in litt. n.”], 436 [as “ Peuhi Mcq. Type. (Bigt.) M.C. ( Ochromyia ) Auchmeromyia Br.Bgst. Senegal.”]. Nomen nudum.

Note. The name as introduced by Brauer & Bergenstamm was possibly based on a specimen labelled peuhi by Macquart [an unpublished Macquart name] from Senegal in NMW (M.C. = museum collection). The expression “ Peuhi Mcq. Type. (Bigt.)” seems to indicate that such a specimen was present in the Bigot collection, of which many specimens were lent out to Brauer (cf. Brauer 1897, 1898, 1899) from Verrall, who had acquired Bigot’s collection in 1893 (A.C. Pont, pers. comm.). However, according to the handwritten list (in OUMNH) that Verrall made of the contents of the Bigot collection at the time of the acquisition, there is no mention anywhere of a specimen named “peuhi” in the collection (A. C. Pont, pers. comm; I have also seen a copy of parts of the pdf file made from Verrall’s list). Perhaps the specimen is in NMW. Whatever the case, Brauer & Bergenstamm provided no description or indication in words on how to distinguish the species; hence their name is a nomen nudum.

Bengalia peuhi Villeneuve, 1914: 253 . Lectotype male (BMNH, examined), by present designation. Type locality: Zambia, Mazabuka.

Note. I have discussed possible syntypes and the lectotype designation below, under Type material.

Bengalia (Ochromyia) peuhi: Malloch 1927: 408 . Burkina Faso (as Haute Volta), Djibouti (locality of Obok), Ethiopia (as Abyssinia), Ghana (as Gold Coast), Kenia (as B.E.A.), Malawi (as Nyasaland).

Bengalia peuhi: Séguy 1933: 79 . Mozambique.

Bengalia bekilyana Séguy, 1935: 132 . Lectotype male (MNHN, examined) by present designation. Type locality: Madagascar (Bekily) (MNHN). For details, see below under Type material.

Bengalia peuhi: Zumpt 1956: 175 . Botswana (as Bechuanaland), Ethiopia (as Abyssinia), Ghana (as Gold Coast), Kenya, Malawi (as Nyasaland), Mozambique (as Port. E. Africa), Namibia (as South West Africa), Nigeria, Somalia (as Somaliland), Sudan, Zambia (as South Rhodesia), South Africa (as Transvaal).

Bengalia peuhi: Séguy 1958: 153 . Burkina Faso (as Haute Volta), Djibouti (as Obock), Ethiopia (as Abyssinie), Mozambique.

Bengalia peuhi: Zumpt 1962a: 65 . Madagascar.

Bengalia bekilyana: Zumpt 1962a: 65 , as synonym of Bengalia peuhi .

Bengalia peuhi: Pont 1980: 791 . Catalogue entry.

Afridigalia elgonia Lehrer, 2005: 34 . Holotype male (BMNH, not examined), by original designation. Type locality: Uganda.

Note. Synonym according to Rognes, 2006: 466.

Afridigalia olapana Lehrer, 2005: 60 . Holotype male (TAU, not examined), by original designation. Type locality: Kenya (TAU).

Note. Synonym according to Rognes, 2006: 466.

Afridigalia arawakia Lehrer, 2006: 2 . Holotype male (BMNH, not examined) by original designation. Type locality: Unknown. Note. Synonym according to Rognes, 2009: 15. The holotype is labelled as being from Kew Park, Jamaica, but is very likely a mislabelled specimen from the Old World tropics (Nigel Wyatt, pers. comm.).

Afridigalia peuhi: Lehrer 2005: 61 . Kenya, Nigeria, South Africa.

Afridigalia peuhi: Lehrer 2006: 9 . Madagascar, Malawi (as Nyassaland), Nigeria, Senegal, South Africa, Sudan, Tanzania, Togo, Zimbabwe.

Afridigalia elgonia: Lehrer 2006: 7 . Kenya, Sudan. Lehrer also reported a male from “ Japon ” (“Place Soba, … coll. Sanousi”), but that is an error. The specimen, which is in BMNH, is labelled “Place: Soba / Host: Termite tunnels / Date: 23.8.76 / Coll. Sanousi” with no mention of Japan. The collector may be Prof. Sulieman Mohamed El Sanousi from the University of Khartoum, so the specimen is likely to be from Sudan (Nigel Wyatt, pers. comm.).

Afridigalia olapana: Lehrer 2006: 9 . Gambia, Kenya, Malawi (as Nyassaland), Nigeria, Sudan, Tanzania, Togo, Zimbabwe.

Bengalia peuhi: Kurahashi & Kirk-Spriggs 2006: 62–63 . Namibia. I have examined all the listed material. For details, see under Material examined, NMNW, below.

Bengalia gaillardi: Kurahashi & Kirk-Spriggs 2006: 62 . Female specimen from Kaross ( Namibia) only. Misidentification, not gaillardi Surcouf & Guyon.

Bengalia spinifemorata: Kurahashi & Kirk-Spriggs 2006: 63 . Six males and eleven females from Uigaranm , Sonderwater , Upper Hungorob ravine, Upper Hungorob , Hungorob ravine, Regenstein ( Namibia) only. Misidentifications, not spinifemorata Villeneuve.

Diagnosis. Male. Length: 10–13.5mm (n=11). Frons at vertex / head width ratio: 0.31–0.34 (mean 0.32, n=14). A pale species recognisable among B. peuhi subgroup members by the shape of the ST5 flap. In most countries the epandrium is shining black, but almost all males from Namibia have a yellow epandrium.

Lunula bare. Fore tibia with a group 3–7 densely set strong spine-like setae, the longest setae are the ones distalmost in the row, longest seta varying from about half as long as width of tibia to as long as width of tibia; these setae much smaller in some males in a series from Madagascar, almost minute; other males from Madagascar have strong fore tibial spine-like setae; the setae always sit on a longish slightly swollen base. Hind tibia with 1–3 pd setae in proximal half, the uppermost weak. Basitarsus with an anteroventral fringe of long setae, longer than diameter of the basitarsus ( Fig. 23) (cf. Villeneuve 1914: 254, “protarse”), a feature shared with B. unicolor (cf. Rognes 2009: 52). Fringes present posteroventrally on the distal half of fore tibia, anteroventrally, ventrally and posteroventrally on distal half of mid tibia, and anteroventrally, ventrally and posteroventrally on distal three fourths of hind tibia.

T5 with a single pair of discal setae (in series of males from Madagascar), or with a more or less regular transverse row of 3–7 discal setae (seen in mainland Afrotropical material). The lectotype has a transverse row of 6–7 discals. According to Lehrer’s description of his Afridigalia elgonia from Kenya and Uganda it has a transverse row of 6 discal setae on T5. Lehrer’s Afridigalia olapana from Kenya has only single pair of discals on T5. Margins of abdominal tergites without or only with small traces of darkening; some specimens have very broad bands of darkening, about 1/3–2/5 of the segment length and very dark on T3–T5, slightly narrower on T1–2 (males from Madagascar). T 5 in one exceptional case without discal setae altogether, see below under specimen from Malawi in BMNH (Davey leg.). Epandrium shining black or dark brown (sometimes yellow in material from Namibia) .

ST5 flap, longer than broad, with roughly parallel, but slightly concave lateral edges, these sometimes diverging somewhat towards the base, and a slightly concave hind edge, the latter sometimes with a narrower notch in middle.

Cerci long and curved. No medial projection of lower bacilliform sclerite (cf. B. minor ).

Distiphallus with dorsolateral wings invisible in dorsal or ventral view. Lateral finger projecting to about outer limit of distal hypophallic lobe as seen in ventral view. In specimen illustrated ( Fig. 30) it projects beyond this limit because of the weakly projecting distal part of external hypophallic lobe. Distal part of external hypophallic lobe usually at level with or projecting beyond outer limit of proximal part in ventral view. The illustrated specimen ( Fig. 30) has an unusually weakly projecting distal part of the hypophallic lobe. Antlers curving upwards, distally with a few tines ( Fig. 32); in some specimens the tip is provided with numerous exceedingly small irregularities, very difficult to make out, even when examined in a microscope with high magnification. Lowermost tine on antler about one third from tip.

Female. Length: 9–13mm (n=10). Frons at vertex / head width ratio: 0.31–0.35 (mean 0.33, n=10). T5 with only 2 discal setae in most female specimens from Madagascar and a few small erect laterodiscals far on each side; in mainland Afrotropical specimens usually with a more or regular transverse row of 4–12 discals, the lateral discals irregularly placed. ST2–ST4 broader than long; ST3 half as long as ST2. ST2 with a group of 2–3 widely separated lateral marginals on each side, those beside it on its outside weaker, but usually no setae between them. ST3 with about 6 similar marginals, lateral edge also with black setae halfway towards middle. ST4 also with about 6–8 marginals and with small setae along whole lateral edge. ST5 triangular, with a rounded hind end, with small black setae along entire margin, 2 stronger setae distally. Disc of all sternites clothed with pale ground setulae. T6 reduced to sclerotisations around spiracles 6 and 7. T 7 in form of elongate rods. T 8 in form of broad plates, almost square in lateral view. ST6 almost completely divided by an oval unsclerotised area in distal three fourths. ST7 completely divided into two sclerites, narrow basally and broad distally. ST8 with diverging parts proximally, distally a single upturned vertical part with numerous setae. Epiproct and cerci straight, short and with soft setae only, no spinous setae as in B. gaillardi . Distal part of vaginal tube with a strong sclerotisation (v.s.) in the lateral wall on right side.

Discussion. Séguy (1935) described the ST5 flap (“la pièce apicale” of the “[c]inquième sternite”) of B. bekilyana as “trapézienne” opposite its condition in B. peuhi , described as “subquadrangulaire à l’apex”. His drawings show the ST5 flap of B. peuhi to have parallel sides and a slightly concave distal margin, whereas the ST5 flap in B. bekilyana is shown with a much narrower base than the concave distal margin. Close examination of the ST5 flap in the dissected lectotype of B. bekilyana with a compound microscope reveals that the lateral edges are parallel, but their basal halves are very sparsely microtrichiose or even without microtrichiae, making the flap appear, under a stereomicroscope, narrower at base than distally where the microtrichiae cover the flap to a much stronger degree. This has obviously misled Séguy into drawing the flap rather narrow at base, thus “trapézienne”.

Lehrer’s (2005, 2006) Afridigalia elgonia , A. olapana and A. arawakia (latter not even diagnosed, but accepted by him as Neotropical taxon) are based on very slight variations in the detailed shape of the ST5 flap. I do not accept these differences as sufficient for distinguishing separate species. Regarding the antler of the distiphallus, Lehrer (2005: 35 fig. 12C [ elgonia ]; 61 fig. 25C [ olapana ]; and 63 fig. 26C [ peuhi ]) and Lehrer (2006: 3 fig. 1C [ arawakia ]) figure the antler tip as terminating in a kind of small narrow “brush” composed of numerous small hair-like projections. I misunderstood the drawings to the effect that the small points (or more exactly, the numerous small strokes of the pen) making up the “brush” were meant to represent denticles (hence my term “denticulate”, cited above), i.e., microscopic structures similar to the ones that cover the lateral finger for example in B. fani Feng & Wei in Feng et al., 1998 ( Rognes 2009: 32 fig. 57; 2011a: 25 fig. 60). After having examined several B. peuhi specimens first hand, I can state with confidence that this is absolutely not the case. They are not denticles or hairs, but simply minute projecting differentiations of the external membrane of antler tip, just like, though much smaller, than the ones in e.g. Bengalia emarginata Malloch or B. emarginatoides Rognes figured in Rognes (2009: figs. 17, 28).

One male from Malawi in BMNH lacks discal setae on the T5 altogether (more details on the specimen can be found below under Material examined). This is an extremely interesting observation, and made me decide that the female holotype of B. africana , also lacking T5 discals, most likely is an aberrant individual of a very commonly collected species .

Biology. The material I have examined, including all the material from Namibia published by Kurahashi & Kirk-Spriggs (2006) and revised by me, has been captured in all the months of the year.

Distribution. Botswana, Burkina Faso, Djibouti, Ethiopia, Gambia, * Ghana, Kenya, * Madagascar, * Malawi, Mozambique, * Namibia, Nigeria, * Senegal, Somalia, * South Africa, * Sudan, Tanzania, Togo, Uganda, * Zambia, Zimbabwe. Also reported from Mali and Yemen by Kurahashi & Kirk-Spriggs (2006) and from Oman, United Arab Emirates and Yemen by Deeming (1996, 2008) [and, doubtfully, from Japan and Jamaica by Lehrer (2006)].

Material examined. Type material. Bengalia peuhi Villeneuve, 1914: 253 . Villeneuve described B. peuhi on the basis of an unspecified number of male and female specimens from ”Transvaal; N.E. Rhodesia; Nigeria; Afrique orientale anglaise; Somaliland, Août-janvier” among material sent to him from the Imperial Bureau of Entomology. He headed the peuhi entry as “ B. Peuhi B. et B. sec. typ.” thus referring to Brauer & Bergenstamm (1891). According to what the Code says about bibliographic reference (Article 72.4.1 of ICZN 1999), the material before Brauer & Bergenstamm when they published the name peuhi should also be reckoned as belonging to the type series of B. peuhi Villeneuve , even though peuhi Brauer & Bergenstamm, 1891 is a nomen nudum. I have found possibly syntypic Villeneuve material in BMNH and OUMNH. One syntype in BMNH is here designated as lectotype (see below) to fix the interpretation of the name. There are six specimens (4 males and 2 females) in OUMNH (ex coll. Bigot, under “ Bengalia depressa ”) that have been identified by Villeneuve as B. peuhi [“ Bengalia … Peuhi BB. sec. Typ.”] (more on all the specimens once under depressa below under treatment of B. depressa ). Of these, 3 males and 1 female are labelled to be from Senegal and 1 male labelled to be from Khartoum [ Sudan] (the second female has no locality label). None of these localities are mentioned in Villeneuve’s paper and no specimens carry labels indicating their original provenance from the Imperial Bureau of Entomology. Neither can I find any mention in Brauer & Bergenstamm (1891) under peuhi that they studied specimens originally placed under depressa in Bigot’s collection. The six specimens have labels reading “ Ph. depressa / EX. COLL. BIGOT” (the first line is handwritten) indicating that Bigot had placed them under that name in his collection. Verrall’s handwritten list (in OUMNH) of the contents of the Bigot collection, made when he acquired it, lists 18 specimens under “ Phumosia depressa ” and 1 specimen under “ Bengalia depressa ”. None of them are marked as having been sent to Brauer, opposite the case with numerous other species in the list. This means that these specimens were never seen by Brauer & Bergenstamm. In addition, these OUMNH specimens are certainly not syntypes, since none of the localities are mentioned in Villeneuve’s 1914 paper. The specimen which is possibly labelled with a Macquart manuscript name and which is cited by Brauer & Bergenstamm under Auchmeromyia peuhi (more details in the first entry in the synonymy, above) may be lost or is still present in NMW. I have not examined it. Lectotype male, by present designation, in BMNH labelled (1) SYN- / TYPE [printed on circular label with a light blue rim]; (2) 12/12/13 // Mazabuka // N.W. Rhodesia // 3400 (R.C.W.) // In house [handwritten on circular cardboard label]; (3) Bengalia / D r Villeneuve det. / Peuhi BB / sec.Typ. [handwritten in Villeneuve’s hand, except line 2 which is printed, on blue label]; (4) Det. Dr. Villeneuve / Pres. by / Impl.Bur.Ent. / 1919—108 [printed]. The locality is in the Southern province of Zambia, quite near the centre of the old Rhodesia, covering the combined area of present day Zimbabwe and Zambia. According to label (4) the specimen originally belonged to the Imperial Bureau of Entomology and was subsequently presented to the BMNH in 1919. Even though Villeneuve writes “N. E. Rhodesia ” and the label reads “N.W. Rhodesia ” I accept it as a syntype. I have labelled it and here designate it as lectotype of Bengalia peuhi Villeneuve in order to fix the interpretation of the name. The specimen, staged on a piece of yellow celluloid, fits the description. All legs are present but the tarsi are lacking on both mid legs and the distal four tarsomeres are lacking on the left hind leg. The darkened hind margins of the abdominal tergites are very faint. The ST5 flap is very similar to the one shown in Fig. 34. I have not dissected it. I have not made any attempt at tracing the remaining syntypes.

Bengalia bekilyana Séguy, 1935: 132 . According to Séguy’s text he had both males and females before him when describing the species, thus syntypes, but he did not indicate how many specimens of either sex. Neither did he use the word “Type”, or “ Holotype ” or similar expression to indicate selection of a holotype. In MNHN there are two specimens under Bengalia bekilyana , one of each sex. Both fit the description. In order to fix the identity of the name, I have labelled and here designate the male as lectotype and the female as paralectotype of Bengalia bekilyana Séguy. The dates are very difficult to decipher from the handwriting on the labels. Séguy gives the capture dates as “…février-avril 1932 …”, but as I read the labels the dates are March and January (see below). Zumpt (1962a: 65) claims to have seen “the type series consisting of 15 specimens ” of Bengalia bekilyana Séguy. I received only two specimens from MNHN when asking for loan of the types. The number 15 may be based on some misunderstanding. Lectotype male, by present designation, in MNHN, labelled (1) TYPE [black print on red label], (2) Madagascar [printed] / Bekily [handwritten] / III 30 [?, handwriting almost illegible] / A. Seyrig [printed]; (3) Bengalia ♂ / bekilyana / type / E. SEGUY det. 1935 [mostly handwritten by Séguy, but last line is printed except 35]. Dissected by KR. Dried T1–5 glued to on card above labels, genitalia in glycerol in glass microvial below labels. The abdomen is yellow with distinct but narrow black margins and shiny black epandrium. T5 with a pair of strong discal setae widely apart. The ST5 flap is typical of peuhi Villeneuve. Paralectotype. MNHN: 1 female labelled (1) COTYPE [red print on yellowish label]; (2) Bengalia ♀ / bekilyana / cotype / E. SEGUY det. 1935 [mostly handwritten by Séguy, but last line is printed except 35]; (3) Madagascar [printed] / Bekily [handwritten] / I. 32. [handwritten] / A. Seyrig [printed]. The female is a pale specimen in good condition, all legs intact.

Other material. BMNH: 1 male labelled (1) MARIMBA / In dambo [a type of African wetland] 3 / miles from lake / Dr. J.B.Davey / 16.1.10 / J.B.D [indistinct handwriting in pencil, line 4 with ink]; (2) Bengalia / depressa Walk / Villeneuve det. [handwritten in ink, but not by Villeneuve]; (3) Pres.by / Comm.Inst.Ent. / B.M.1953-343 [printed]; (4) AFRICANA [handwritten in blue with ball-point pen on white label]; (5) Bengalia (m) / peuhi Villeneuve, 1914 / K. Rognes det. 2012 [printed on white label]. The specimen was placed in BMNH under B. depressa . The specimen has the abdominal tip and ST5 flap removed from below and carried a large plastic vial with dissected parts in glycerol. There were three items in the vial: ST5 flap and parts of ST5; epandrium, cerci and parts of surstyli and bacilliform sclerites; hypandrium with pre- and postgonites and a broken aedeagus. These were transferred by KR to a glass microvial now pinned below label (4). I have added my determination label as B. peuhi . Left fore and mid leg present, other legs lost. The specimen lacks any discal setae on T5. Lehrer appears to have dissected it to judge from the type of removal of genitalia from the abdominal tip and the big plastic vial, but he did not identify or label the specimen. The dissection was unsuccessful, for the distal part of the distiphallus is absent (destroyed). The shape of the intact ST5 flap, the pale only anepimeral ground vestiture, the very short and stubby pv setae all along the mid femur and the spine-like setae on fore tibia clearly identify the specimen as B. peuhi . The label reading “AFRICANA” must have been put there by someone (Lehrer?) having observed the lack of discal setae on the T5, and thinking the specimen might be the male of the nominal species B. africana . However, the anepimeron is clothed only with pale setulae, opposite the condition in B. africana where all these setulae are black. The alleged determination by Villeneuve as B. depressa seems mysterious. I do not accept this specimen as one of the syntypes of B. unicalcarata , since Villeneuve hardly would have thought of this specimen as belonging to that taxon. J. B. Davey was a medical officer in British Central Africa (later Nyasaland, now = Malawi) and was a participant of the Royal Society’s Sleeping Sickness Commission during 1910–11. Most of his time was spent collecting tsetse flies on the shores of Lake Nyasa. His diary has entries for 15 and 18 January 1910, i.e., the day before and two days after he captured the aberrant B. peuhi ( Vaughan 1991: 36–37) . The locality Marimba is in present day Malawi, but I cannot find it on any map or in Google Earth. However, a Google search for “Marimba Malawi”, results in hits concerning a Kota-Kota Meteorite, Malawi. The locality is now spelt Nkhotakota and is in Central Malawi. BMSA: South Africa [5 males 2 females]: 1 male labelled (1) SIBASA / SE 22 30 Cd / 7. VII. 1980 / J. E. CRAFFORD / DEPT. ENTOMOLOGY / UNIVERSITY OF PRETORIA (handwritten on white label, except last two lines which are printed]; (2) Ex. Dept. Entomology / University of Pretoria Coll. / Donated 2009 [printed on yellow label]; (3) Entomology Dept. / National Museum / P.O. Box 266 / Bloemfontein 9300 / South Africa [printed on blue label]; (4) BMSA (D) / 18168 [printed on white label, printed side downwards]. T5 with four evenly spaced discal setae in a row, erect ground setulae laterally and ventrally on T5. 1 male labelled (1) SUIDAFRIKA / Noordelike P. / Tzanee / 23°50'S 30°10'E / 13.III.1999 / C.E.VENTER [printed on white label]; (2) Ex. Dept. Entomology / University of Pretoria Coll. / Donated 2009 [printed on yellow label]; (3) Entomology Dept. / National Museum / P.O. Box 266 / Bloemfontein 9300 / South Africa [printed on blue label]; (4) BMSA (D) / 18176 [printed on white label, printed side downwards]. T5 with 2 discal setae only. 1 male labelled (1) C. J. COLINJ / ACORNHOEK / APRIL 1971; (2) Ex. Dept. Entomology / University of Pretoria Coll. / Donated 2009 [printed on yellow label]; (3) Entomology Dept. / National Museum / P.O. Box 266 / Bloemfontein 9300 / South Africa [printed on blue label]; (4) BMSA (D) / 18159 [printed on white label, printed side downwards]. T5 with 3 strong discals. 1 male labelled (1) W K R [handwritten] / Marno [? handwritten; difficult to interpret] Tvl. [= Transvaal; printed] / 27.xii.1929 [handwritten, except the number 19]; (2) Ex. Dept. Entomology / University of Pretoria Coll. / Donated 2009 [printed on yellow label]; (3) Entomology Dept. / National Museum / P.O. Box 266 / Bloemfontein 9300 / South Africa [printed on blue label]; (4) BMSA (D) / 18167 [printed on white label, printed side downwards]. T5 with 2 discals only. 1 male labelled (1) SOUTH AFRICA [printed]: T.v.L. / S E 26°20' 28°46' / Dornd’ooi / 25 - IV – 1988 / M.v. Niekerk / Department of Entomology [printed] / University of Pretoria [printed] [white label with black rim]; (2) Ex. Dept. Entomology / University of Pretoria Coll. / Donated 2009 [printed on yellow label]; (3) Entomology Dept. / National Museum / P.O. Box 266 / Bloemfontein 9300 / South Africa [printed on blue label]; (4) BMSA (D) / 18170 [printed on white label, printed side downwards]. T5 with row of four discals (the rightmost one lost, only basal pore remains. 1 female labelled (1) E. R. L. / Mei 1971 / Pretoria; (2) Ex. Dept. Entomology / University of Pretoria Coll. / Donated 2009 [printed on yellow label]; (3) Entomology Dept. / National Museum / P.O. Box 266 / Bloemfontein 9300 / South Africa [printed on blue label]; (4) BMSA (D) / 18178 [printed on white label, printed side downwards]. T5 with row of eight strong, several additional short discal setae. 1 female labelled (1) E. R. L. / Mei 1971 / Pretoria; (2) Ex. Dept. Entomology / University of Pretoria Coll. / Donated 2009 [printed on yellow label]; (3) Entomology Dept. / National Museum / P.O. Box 266 / Bloemfontein 9300 / South Africa [printed on blue label]; (4) BMSA (D) / 18180 [printed on white label, printed side downwards]. T5 with 2 strong discals, plus at least 3 additional small ones on each side. MNHN: Madagascar (12 males, 10 females): 6 males and 4 females labelled: (1) Madagascar Sud-Ouest / SEPT LACS 50m / Tuléar / VI–VII 57 Andria R. [photographic paper label]; (2) INSTITUT / SCIENTIFIC / MADAGASCAR [printed black on pale green label]. One male and one female have been dissected by KR and have the dried T1–5 glued to piece of card above labels, and the genitalia in glycerol in glass microvial pinned below labels. 1 male and 1 female labelled: (1) Amplijeroa 170m / Ankarafantsika / –I–57 R.E. [printed label]; (2) INSTITUT / SCIENTIFIC / MADAGASCAR [printed black on pale green label]. 1 male labelled (1) Madagascar Sud-Ouest / Tuléar-Sakaraha / Zombitsy 630m / Raharitzonina [printed label]; (2) INSTITUT / SCIENTIFIC / MADAGASCAR [printed black on pale green label]. 4 males and 2 females labelled (1) Andobo 190m / forêt Antsingy / dct Antsalova / –II– 57 P. Griv. [printed label]; (2) INSTITUT / SCIENTIFIC / MADAGASCAR [printed black on pale green label]. One of the males has been dissected by KR and has the dried T1–5 glued to piece of card above labels, and the genitalia in glycerol in glass microvial pinned below labels. 3 females labelled (1) Madagascar Sud / Poste Adm. de Bsivory / Marotsiraka / XI–59 Randriamasy [handwritten in black ink]; (2) INSTITUT / SCIENTIFIC / MADAGASCAR [printed black on pale green label]. One female has been dissected by KR. The dried T1–5 glued to piece of card above labels, and the genitalia in glycerol in glass microvial pinned below labels. The ovipositor has been slit open between tergites and sternites on one side. Senegal: 1 female labelled (1) Muséum Paris / SENEGAL / M’Bour / 10.XI.1980 / B. SIGWALT [blue label, first line printed, others handwritten]; (2) Piège de / Malaise / de jour [printed on white label]; (3) My determination label. Sudan: 1 male labelled (1) KOGONI / SOUDAN / 24.X.55 / J.-G.POINTEL [printed, except date which is hand-written over lines 1 and 2]. (2) My determination label. The specimen has 6 strong discal setae in a transverse row on T5.? locality: 1 female labelled (1) Bol – Juin 1980 / Dr. Gaillard [handwritten]; (2) Bengalia depressa Walk [handwritten]; (3) KR’s determination label. No locality label, unless “Bol” means a locality. NMNW: Namibia [23 males and 33 females (correctly identified as B. peuhi ); 1 female from Kaross (misidentified as B. gaillardi ); 6 males and 11 females from Uigaranm, Sonderwater, Upper Hungorob ravine, Upper Hungorob, Hungorob ravine, Regenstein (misidentified as B. spinifemorata by Kurahashi & Kirk-Spriggs 2006: 63)]. The details regarding these specimens are not repeated here; the reader is referred to Kurahashi & Kirk-Spriggs (2006). OUMNH [4 males, 2 females, all from the Bigot collection under Bengalia depressa (Walker) , re-identified by Villeneuve and now placed under a single blue handwritten drawer label reading “ Bengalia / D r Villeneuve det. [this line printed] / Peuhi BB. / sec. Typ.” in Villeneuve’s hand] (only one of these specimens, the one dissected, has been given my determination label): Senegal: 1 male labelled (1) TYPE [printed text in red on round white label; (2) Senegal [black ink handwriting]; (3) Ph. depressa [black ink handwriting] / EX. COLL. BIG. [printed in black]; (4) Oxford University / Museum Natural / History ( OUMNH) [printed in black]. 1 male labelled (1) Senegal [black ink handwriting]; (2) Ph. depressa [black ink handwriting] / EX. COLL. BIG. [printed in black]; (3) Oxford University / Museum Natural / History ( OUMNH) [printed in black]. 1 male labelled (1) Senegal [black ink handwriting]; (2) Ph. depressa [black ink handwriting] / EX. COLL. BIG. [printed in black]; (3) Oxford University / Museum Natural / History ( OUMNH) [printed in black]; (4) Bengalia (m) / peuhi Villeneuve, 1914 / K. Rognes det. 2011 [printed]. Dissected by KR. Area of upper lip between two strengthenings more sclerotised than the rest, appearing as a broad sclerotisation at middle. 5 exceedingly small tines distally on antlers. Very strongly sclerotised genitalia. No vestiture on underside of distal surstylus. 1 female labelled (1) Senegal [black ink handwriting]; (2) Ph. depressa [black ink handwriting] / EX. COLL. BIG. [printed in black]; (3) Oxford University / Museum Natural / History ( OUMNH) [printed in black]. Sudan: 1 male labelled (1) Khartoum [black ink handwriting]; (2) Ph. depressa [black ink handwriting] / EX. COLL. BIG. [printed in black]; (3) Oxford University / Museum Natural / History ( OUMNH) [printed in black].? locality: 1 female labelled (1) Ph. depressa [black ink handwriting] / EX. COLL. BIG. [printed in black]; (2) Oxford University / Museum Natural / History ( OUMNH) [printed in black]. No locality label. ZMUN: Ghana: 2 females labelled: (1) GOLD COAST [Ghana] / N. TERRITORIES / KOFABA / FEB. 1916 / DR J J SIMPSON [printed]; (2) Department of Zoology / Natural History Museum / University of Oslo / ( ZMUN) / World collection [printed]. One of the females carries a handwritten determination label reading (3) Bengalia / peuhi B.B. I have added my determination label to both specimens. Both females have a transverse row of 9 discal setae on T5 (4 on the left, 5 on the right side).