Agathidium akrogeneios Miller and Wheeler, 2005

MILLER, KELLY B. & WHEELER, QUENTIN D., 2005, Slime-Mold Beetles Of The Genus Agathidium Panzer In North And Central America, Part Ii. Coleoptera: Leiodidae, Bulletin of the American Museum of Natural History 2005 (291), pp. 1-167 : 133-134

publication ID	https://doi.org/ 10.1206/0003-0090(2005)291<0001:SBOTGA>2.0.CO;2
persistent identifier	https://treatment.plazi.org/id/038387B3-37C9-B981-FD72-57D7FCFC0A0D
treatment provided by	Felipe
scientific name	Agathidium akrogeneios Miller and Wheeler
status	sp. nov.

Treatment

Agathidium akrogeneios Miller and Wheeler View in CoL , new species Figures 158 View Figs , 309, 310 View Figs , 369 View Fig

TYPE MATERIAL: Holotype, Ƌ in CMNC labeled ‘‘GA: Dade Co., Cloud land Canyon St. Prk. 16.v.1972, S.&J. Peck Rhodo. Litter,B236/ HOLOTYPE Agathidium akrogeneios Miller and Wheeler, 2003 [red label with black line border]’’.

TYPE LOCALITY: United States, Georgia, Dade Co., Cloudland Canyon State Park.

DIAGNOSIS: This species is similar to other A. dentigerum subgroup members that have a large, acute, somewhat falcate male metafemoral tooth (fig. 158), very narrow metasternum with the oblique metasternal carinae relatively prominent and meeting medially in a large, posteriorly directed triangular lobe, large male metasternal fovea, and somewhat reduced eyes. However, this species has a very prominent lobelike tubercle medially on the gula, making it most similar to A. stephani and A. dentigerum . From these species it may be distinguished by the shape of the median lobe, which is straight in lateral aspect from near the base to near the apex where it is abruptly recurved dorsally (fig. 310).

DESCRIPTION: Body small (TBL = 2.36– 2.45 mm), rounded, robust (PNW/TBL = 0.45–0.51), strongly contractile.

Head, pronotum, and elytra red; venter red to yellowbrown; antennae, palpi, and legs yellow to yellowbrown.

Head broad (MDL/OHW = 0.54–0.62), dorsal surface flattened, dorsoventrally compressed; with very fine punctures, each with a short, very fine seta, surface between punctures shiny, smooth; frontoclypeal suture obsolete medially; eyes moderately prominent and protruding, but dorsoventrally compressed, largefaceted; gula broad, flattened, with prominent median lobelike tubercle; antennomere ratios: length I:II:III = 1.6:1.0: 1.0, width VII:VIII:IX = 1.0:1.0:2.3. Pron otum very large, broad (PNL/PNW = 0.70– 0.76), strongly convex, anterolateral lobes strongly produced, lateral margin broadly curved, not angulate; with very fine, sparse punctures, each with a short, very fine seta, surface between punctures smooth, shiny. Elytra broad, lateral margins strongly rounded, apically rounded (SEL/ELW = 0.86–1.15); punctation similar to pronotum; sutural stria absent. Flight wings strongly reduced. Mesosternum broad, broadly convex; medial carina prominent. Metasternum narrow (MTL/ MTW = 0.10–0.15), slightly concave medially, distinctly sloping dorsad anteriorly; oblique femoral carinae not prominent laterally, medially meeting in very prominent, posteriorly directed triangular flange.

Male tarsi 554; pro and mesobasotarsomeres slightly laterally expanded, with small ventral field of spatulate setae; mandibles not modified; metafemur slender, with very sharp, acuminate, large tooth subapically on posterior margin (fig. 158); metasternal fovea posterior, prominent, round with large brush of long, fine setae. Median lobe in lateral aspect slender, basally bent, straight thereafter, apical portion tapered, straight, apex abruptly and sharply recurved dorsally (fig. 310); in ventral aspect moderately broad, lateral margins broadly curved, apex abruptly narrowed and with medial, narrowly rounded prominence (fig. 309); operculum divided into two long, slender rami, apices slightly divergent and each narrowly rounded (fig. 309); lateral lobes moderately broad, evenly curved in lateral aspect, apex pointed or narrowly rounded with 2 long subapical setae (fig. 310).

Female tarsi 544.

ETYMOLOGY: This species is named akrogeneios from the Greek word meaning ‘‘with a prominent chin’’, for the prominent gular tubercle in this species.

DISTRIBUTION: This species is found in southeastern North America from Texas to Georgia and north to Virginia (fig. 369).

PARATYPES: UNITED STATES: Alabama: Jackson Co.: 5 mi NW Princeton , 19 May 1972, forest litter, S Peck (1, PECK) ; 5 mi N Garth , 19 May 1972, S Peck (8, PECK) ; Madison Co.: Monte Sano St. Park, 17 May 1972, S and J Peck (3, PECK). Georgia: Dade Co. : Cloudland Canyon St. Park., 16 May 1972, Rhododendron litter, S and J Peck (2, PECK). Kentucky: Edmonson Co. : Mammoth Cave Natl Park, Doyle Valley , 30 Aug 1969, log litter, S Peck, A Fiske (1, PECK). Louisiana: W Feliciana Co. : Magnolia Glen TNC property near Tunica WMA, 16 Nov 1996, mixed mesophytic, Berlese, CE Carlton (24, LSAM) ; Tunica WMA Magnolia Glen TNC Property , 10 mi NW St Francisville, 25 Mar 1998, mesophytic forest, Berlese, CE Carlton (19, LSAM) ; Cabin area 30°47̍N, 91°15̎W, 1 Feb 2001, pitfall, AR Cline (3, LSAM). Tennessee: Hardin Co.: 0.5 mi W Olive Hille, 28 May 1991, old growth, Berlese, C Carlton (3, LSAM). Texas: Sabine Co. : 9 mi E Hemphill, 11 May 1988, beech, magnolia litter, Berlese, R Anderson (2, LSAM). Virginia: Lee Co. : Cumberland Gap Natl Park. nr Skyline Cave , 10 Jul 1971, dry litter at log, W Shear (1, AMNH) .

DISCUSSION: This species has been collect ed from February to November. It has been found in a variety of habitats, including ‘‘mesophytic forest’’, and various litter types, including logs, Rhododendron , beech, and magnolia.