Pyxine schmidtii Vain.

Mongkolsuk, Pachara, Meesim, Sanya, Poengsungnoen, Vasun & Kalb, Klaus, 2012, The lichen family Physciaceae in Thailand-I. The genus Pyxine, Phytotaxa 59 (1), pp. 32-54 : 50

17. Pyxine schmidtii Vain. , Hedwigia 46: 170 (1907).

Holotype:— THAILAND. Trat; In insula Koh Chang prope Lem Dan , ad truncos arborum in silva, 1900, J . Schmidt 25 ( TUR-VAIN. 8695!)— Pyxine papuana Kashiw., Bull. Natl Sci. Mus. , Tokyo, B , 3 (2): 68 (1977). Holotype: PAPUA NEW GUINEA. Morobe District; Mt. Kaindi, Wau , 1300 m, H . Kashiwadani 10658 ( TNS!) .

Thallus 4–10 cm wide, adnate, subdichotomously lobate. Lobes radiating, discrete or rarely contiguous and imbricate, plane to convex but often slightly concave towards the tips, 0.5–1.2 mm wide, subrotund at the apices. Upper surface white to whitish grey, pruinose, especially towards the lobe tips; polysidiangia, isidia and soralia absent. Pseudocyphellae marginal and laminal, irregular to linear or becoming reticulate. Medulla uniformly white. Lower surface black in the centre, paler towards the margin; rhizines ± dense, furcate. Apothecia common, obscurascens - type, 0.5–2.0 mm wide; disc epruinose. Internal stipe distinct, white to brown, K-, P-. Ascospores 16–20 × 6–8 µm. Pycnidia not seen.

Chemistry: Cortex K + yellow, C-, KC-, P + yellow, UV-; medulla K-, C-, P + orange-red; containing atranorin (minor), chloroatranorin (minor), testacein (major), unknown terpenes (minor).

Distribution and habitat:—Rare on bark in dry evergreen, lower montane rainforest, lower montane scrub and mixed deciduous forests. Also in SE Asia, Australia and Papua New Guinea.

Notes:—This species is characterized by the adnate thalli, the absence of polysidiangia, soredia, isidia and norstictic acid and the presence of cortical atranorin, medullary testacein and obscurascens - type apothecia.

Material from Thailand examined:— Loei; Phu Luang Wildlife Sanctuary, Huai Nam San, on the bark of an unidentified tree in mixed deciduous forest, 787 m, 17°20'36'' N, 101°30'17'' E, 11 November 2008, S . Meesim & K . Buaruang MS 0142 ( RAMK).— Loei; Phu Luang Wildlife Sanctuary, Khok Prommachan , on rocks in lower montane scrub, 1487 m, 17°16'53'' N, 101°31'22'' E, 26 June 2008, S . Meesim MS 41 ( RAMK).— Loei; Phu Luang Wildlife Sanctuary, Khok Huai Toei , on rocks in lower montane rainforest, 1240 m, 17°19'46'' N, 101°31'26'' E, 27 June 2008, S . Meesim 57, 58 ( RAMK).— Nakhon Ratchasima; Khao Yai National Park , on the bark of an unidentified tree in dry evergreen forest, 850 m, 14°30' N, 101°37' E, 24 May 2000, W . Polyiam 19575 ( RAMK) .