Trechus tilitshoensis Schmidt, 1994

Schmidt, Joachim, 2009, Taxonomic and biogeographical review of the genus Trechus Clairville, 1806, from the Tibetan Himalaya and the southern central Tibetan Plateau (Coleoptera: Carabidae: Trechini) 2178, Zootaxa 2178 (1), pp. 1-72 : 25-26

Trechus tilitshoensis Schmidt, 1994

( Figs. 30 , 42 )

Catalogue: Trechus tilitshoensis Schmidt, 1994: 130 . Locus typicus: Central Nepal, Manang Distr., Plateau above Tilitshó Lake at N-slope of Annapurna Massif, altitude approximately 5000 m.

Type material: Holotype male, with label data “NEPAL-HIMALAYA, Annapurna-N-Abfall, W-Manang, 6- 8.10.92”, “Plateau über dem Tilitschok-Lake 5000 m, lg. Schmidt”, “ HOLOTYPUS Trechus tilitshoensis des. J. Schmidt 1993” ( SMTD) . Paratypes: 8 males, 3 females, with same label data as holotype ( CSCHM, SMTD) ; 10 males, 7 females, Annapurna Mts. , Tilitshó Lake W Manang, 4950–5200 m, 4.VI.1993, leg. Schmidt ( CSCHM) ; 2 males, Annapurna Mts. , Thorong Pass N Manang, E slope, 4900–5200 m, 8.VI.1993, leg. Schmidt ( CSCHM) .

Additional material: NEPAL: 8 males, 3 females, Annapurna Mts., Manang Distr., E slope Kang La Pass , 5000 m, 3.VI.1994, leg. J. Schmidt ( CSCHM) ; 3 males, 1 female, Annapurna Mts. , Yakkharka N Manang, 4500 m, 28.V.1996, leg. J. Schmidt ( CSCHM) ; 10 males, 2 females, N Annapurna Mts., Gungdang N-slope, W Thorung Phedi, 4600–4900 m, 30.V.1996, leg. J. Schmidt ( CSCHM) ; 24 males, 12 females, Dhaulagiri, upp. Yakkharka [place above Marpha north of Tukuche Peak], 4500–4600 m, 12.7.1998, leg. C. Berndt & J. Schmidt ( CSCHM) .

Identification: See key above.

Relationships: This species and the Western Nepalese species T. aedeagalis sp. n., T. eremita sp. n., T. franzianus Mateu & Deuve, 1979 , T. muguensis sp. n., and T. sculptipennis sp. n., together forming a group of closely related species which, in external morphology, differ very slightly from each other or, in some cases are almost identical, but which evolved remarkable differences in genital morphology. Currently, based on these characters it seems impossible to determine sister species relationships.

Distribution: Fig. 98 . Tibetan Himalaya of Manang and Mustang Districts, Central Nepal. The species is known from several localities north of Annapurna Massif as well from the Northeast slope of Dhaulagiri Himal.

Habitat: Edaphic species of the higher alpine zone; vertical distribution approximately 4900–5200 m. The specimens were found on humid, gently inclined slopes and along small depressions, often close to snow fields and melting water.

Figures

FIGURE 98. Map of central parts of Tibet and the Himalaya showing distribution of species of the Trechus thibetanus group (white circles), of the Trechus pumoensis group (white squares), of the Trechus rolwalingense group (black circles), and of the Trechus stratiotes group (black squares). 1, T. namtsoensis sp. n. 2, T. dongulaensis sp. n. 3, T. thibetanus Jeannel, 1928. 4, T. glabratus sp. n. 5, T. thorongiensis Schmidt, 1994. 6, T. eutrechoides eutrechoides Deuve, 1992. 7, T. eutrechoides mondaensis Deuve, 1997. 8, T. pumoensis Deuve, 1997. 9, T. tilitshoensis Schmidt, 1994. 10, T. franzianus Mateu & Deuve, 1979. 11, T. muguensis sp. n. 12, T. eremita sp. n. 13, T. aedeagalis sp. n. 14, T. sculptipennis sp. n. 15, T. rolwalingense rolwalingense ssp. n. 16, T. rolwalingense daldunglana ssp. n. 17, T. stratiotes stratiotes ssp. n. 18, T. stratiotes malikasthana ssp. n.

FIGURES 28–35. Head, pronotum. Fig. 28, Trechus pumoensis Deuve, 1997, paratype, male. Fig. 29, T. eremita sp. n., paratype, female. Fig. 30, T. tilitshoensis Schmidt, 1994, paratype, male. Fig. 31, T. aedeagalis sp. n., paratype, male. Fig. 32, T. antonini Deuve, 1997, non-type female specimen from Lhachen La. Fig. 33, T. yeti sp. n., paratype, male. Fig. 34, T. cf. damchungensis Deuve, 1997, paratype, male from Tangula Pass. Fig. 35, T. hodeberti Deuve, 1997, non-type female specimen from Lhachen La.

FIGURES 39–51. Aedeagal median lobe with sclerotized portions of internal sac in lateral view (Figs. 39, 40, 42, 43, 45, 47, 49; Figs. 39, 40, 47, 49 in addition with parameres) and in dorsal view (Figs. 41, 44, 46, 48, 50). Fig. 39, Trechus pumoensis Deuve, 1997, paratype. Figs. 40, 41, T. franzianus Mateu & Deuve, 1979, non-type specimen from Dhauli Lake. Fig. 42, T. tilitshoensis Schmidt, 1994, non-type specimen from Gungdang N-slope. Fig. 43, 44, T. muguensis sp. n., paratype (43) and holotype (44). Figs. 45, 46, T. eremita sp. n., paratype. Figs. 47, 48, T. sculptipennis sp. n., paratype. Figs. 49, 50, T. aedeagalis sp. n., paratype. Fig. 51, T. stratiotes sp. n., paratype.