Hebynthus phyrra, Noyes, 2023

Noyes, John Stuart, 2023, ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera), Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) 2 (11), pp. 1-921 : 689-690

Hebynthus phyrra sp.nov.

(Figs 1888-1892)

DIAGNOSIS. Female (length about 1.0mm): antenna (Fig. 1891) with scape pale brown, orange apically; funicle brown, mixed orange-brown; coxae dark brown, legs largely brown; fore wing (Fig 1892) hyaline; head about 3.7X as wide as frontovertex; malar sulcus absent; antenna (Fig. 1891) with scape about 5.6X as long as broad; pedicel nearly as long as F1-F3 combined; all funicle segments transverse, distal segments subquadrate; linear sensilla on F3-F6; sensory part of clava extending about 0.4X along ventral surface; fore wing (Fig. 1892) about 2.3X as long as broad; ventral surface of costal cell with only a single complete line of setae; mid tibial spur about 1.3X as long as basitarsus; ovipositor slightly exserted with exserted part about 0.4X as long as mid tibial spur; gonostylus about as long as mid tibial spur. Male: unknown.

Female (holotype): length, including ovipositor, 1.02mm; excluding ovipositor, 0.96mm ( CPD) .

Head mainly black; frontovertex with a relatively weak dark green and purple sheen; temple and gena with a mixed weak brassy and coppery sheen; area immediately below eye distinctly green, brassy and coppery; area between eye and scrobe green and coppery; interantennal prominence and scrobes with a metallic coppery purple sheen; frontovertex, gena and interantennal prominence with fairly conspicuous dark brown setae; antenna (Fig. 1891) with radicle dark brown; scape pale brown, orange-brown towards apex and internally; pedicel dark brown, orange-brown ventrally and apically; F1-F5 orange-brown, F6 pale orange, clava brown; maxillary palpus white, terminal segment orange; pronotum black with weak coppery and green reflections; propleuron and prosternum dark brown; mesoscutum slightly dull, metallic blue-green with some weak coppery reflections, posterior margin dark brown with a slight coppery sheen; tegula dark brown with a slight brassy sheen; axilla black with a weak coppery lustre; scutellum black and much duller than mesoscutum, but with weak brassy reflections, sides and apex with a slight blue-green sheen, vertical apex and side, very shiny metallic green mixed with some dark blue and coppery; metanotum dark brown; mesopleuron dark brown with a coppery, brassy and purple sheen; mesosternum dark brown; fore coxa dark brown; fore femur dark brown, tibia pale orange with a broad, subbasal, brown band, tarsus pale orange; mid coxa dark brown; mid femur white basally, but mostly dark brown, tibia white to pale orange, brown at base and with a broad, subbasal brown ring, spur and tarsus pale orange; hind coxa dark brown; hind femur dark brown, extreme base white, tibia dark brown, orange at apex, tarsus pale orange-brown; wings (Fig. 1892) hyaline, venation pale orange; propodeum dark brown with a weak brassy sheen, side metallic blue-green anteriorly, mixed green posteriorly, a group of about 8 or 9 scattered, conspicuous, translucent, white setae anterolateral and posterior to spiracle; gaster dark brown, with a fairly strong purple and coppery sheen dorsally, Gt1 contrasting and strongly metallic blue with some green areas, side of gaster metallic green, blue and coppery reflections, venter slightly duller with mixed green, purple and blue, outer plates of ovipositor relatively dull coppery, purple and blue; gonostylus dark brown.

Head about 3.7X as wide as frontovertex, in profile about 1.7X as high as deep, anteriorly fairly evenly curved from occipital margin to top of scrobes and then virtually straight to mouth margin, with interantennal prominence slightly protuberant; occipital margin sharp, carinate; ocelli forming an angle of about 70°; frontovertex slightly shiny, with fairly regular, moderately deep, polygonally reticulate sculpture of mesh size generally subequal to eye facet; frontovertex without setae in front of anterior ocellus, a line of sparse setae along inner eye margin continued along lower eye margin; temple and gena more shiny than frontovertex, with much shallower, longitudinally elongate, strigose to striate-reticulate sculpture; eye reaching occipital margin, virtually naked, but with extremely short, inconspicuous setae that are each much shorter than diameter of facet, eye separated from scrobe by about diameter of ocellus, area between eye and scrobe with shallow, transversely elongate, polygonally reticulate sculpture; scrobes shallow, narrowly ∩-shaped, meeting, with distinct polygonally reticulate sculpture, weakly margined dorsally and laterally; interantennal prominence weak, rounded dorsally, with shallow, polygonally reticulate sculpture; antenna as in Fig. 1891; scape subcylindrical, about 5.6X as long as broad; all funicle segments subquadrate, segments larger and broader distad, linear sensilla present only on F3-F6; clava slightly broadened, nearly as long as funicle, outer suture strongly oblique, sensory area enlarged, extending ventrally about 0.4X length of clava, apex strongly obliquely truncate; malar sulcus absent; clypeal margin hardly emarginate medially; mandible with two acute teeth and a truncate upper tooth. Relative measurements: HW 57.5, HH 56.5, FV 15.5, POL 9.5, OOL 0.5, OCL 2.5, AOL 8.5, EL 37, EW 29.5, MS 23.5, SL 28, SW 5.

Thorax in dorsal view with pronotum short, mostly hidden by head; pronotum dorsally with imbricate-reticulate to polygonally reticulate sculpture that is slightly shallower than sculpture on frontovertex, and of larger mesh; mesoscutum with polygonally reticulate sculpture that is of similar mesh size, but slightly shallower than that on pronotum; axilla with polygonally reticulate sculpture that is slightly deeper than that on pronotum and of slightly larger mesh; dorsal part of scutellum with sculpture similar to that on axilla, but conspicuously deeper and coarser and more regularly polygonal medially to slightly more elongate laterally, vertical apex and side completely smooth and shiny; visible part of mesoscutum about 2.2X as broad as long; scutellum about 1.1X as broad as long; fore wing with venation and distribution of setae as in Figs 1889, 1892; ventral surface of costal cell with only a single complete line of setae; mid tibial spur about 1.3X as long as basitarsus; propodeum medially about 0.1X as long as scutellum, virtually smooth. Relative measurements: FWL 61, FWW 25; HWL 45, HWW 12.

Gaster with hypopygium (Fig. 1890) reaching about 0.6X to apex; syntergum about 0.65X as long as mid tibia, with apex acute; ovipositor (Fig. 1888) slightly exserted, the exserted part about 0.4X as long as mid tibial spur or about 0.16X mid tibia; gonostylus about as long as mid tibial spur. Relative measurements: OL 70, GL 22 [MT 56].

Variation. Only holotype examined.

Male. Unknown.

DISTRIBUTION. Costa Rica.

HOSTS. Unknown.

MATERIAL EXAMINED.

Type material. Holotype E: COSTA RICA, Cartago, PN Tapanti-Macizo de la Muerte, Send. a Rio Humo , LN 188100 560500, 1400-1500m, xii.1999 - i.2000 (M. Alfaro) . Holotype in NHMUK .

COMMENTS. Hebynthus phyrra is probably closest to solex and cleon (see comments under solex , p. 694)