Dalek ananke, Noyes, 2023

Noyes, John Stuart, 2023, ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera), Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) 2 (11), pp. 1-921 : 642-644

Dalek ananke sp.nov.

(Figs 1755-1761; Hab. E 260)

DIAGNOSIS. Female (length about 1.7-1.9mm): scape (Fig. 1757) dark brown, apex orange; mesoscutum bright metallic dark green or blue-green; all coxae dark brown; clypeal margin (Fig. 1761) with a very short, rounded, median lobe; OOL much less than OCL; antenna (Fig. 1757) with scape about 3.3X as long as broad; funicle with F1 smallest, remaining segments subequal, mostly about 1.5X as long as broad, F6 relatively broader; linear sensilla only on F2-F6; mesoscutum with shallower sculpture than scutellum (Fig. 1758); sculpture on scutellum extending to side and apex; mid tibial spur about as long as basitarsus; fore wing (Fig. 1759) about 2.5X as long as broad; propodeum with about 15 setae around spiracle; apex of syntergum acute; ovipositor (Fig. 1756) longer than mid tibia, exserted part about 0.5X as long as mid tibial spur; gonostylus very slightly longer than mid tibial spur. Male: unknown.

Female (holotype): length, including ovipositor, 1.71mm; excluding ovipositor, 1.60mm (CPD).

Head generally black with a coppery purple sheen; ocellar and scrobal areas dull, metallic dark green; temple and gena metallic green near eye, posteriorly coppery and brassy, gena coppery purple near mouth; mouth margin coppery purple; frontovertex, gena and interantennal prominence with inconspicuous, brown setae; antenna (Fig. 1757) with radicle mostly dark brown, mixed with some pale orange; scape mostly dark brown, extreme apex pale orange; pedicel dark brown, pale orange ventrally and at extreme apex; flagellum brown; maxillary palpus white, terminal segment very pale orange; thorax generally dark brown to black; pronotum with a moderate coppery purple and dark green sheen; mesoscutum relatively bright metallic dark green and blue-green, with some coppery purple reflections, posterior margin narrowly coppery purple; tegula dark brown with a coppery purple sheen; axilla dark purple-brown with a coppery purple sheen; scutellum relatively bright metallic dark green and blue-green, with some coppery reflections, extreme apex with a coppery and dark blue sheen; mesoscutum and scutellum clothed with fairly inconspicuous dark brown setae; metanotum dark purple-brown; mesopleuron very dark brown, with a brassy sheen mixed with some coppery purple; mesosternum dark brown; all coxae dark brown; fore femur dark brown with extreme apices pale orange, tibia dark brown with extreme base and apical 0.3X pale orange, tarsus dusky pale orange, apical tarsomere brown; mid femur off-white in proximal 0.2X, apical 0.2X pale orange, otherwise dark brown, tibia dusky pale orange with a broad, brown, subbasal ring extending to about 0.6X, spur and tarsus pale orange, distal tarsomeres slightly dusky, apical tarsomere brown; hind femur dark brown, extreme apex pale orange, tibia mostly dark brown with extreme base and apex pale orange-brown, tarsus dark orange-brown, apical tarsomeres dark brown; wings mostly hyaline, but fore wing (Fig. 1759) with a small, weak, infuscate area immediately below marginal and stigmal veins; propodeum dark brown, with a weak purple and brassy sheen, side with a dull metallic green sheen with about 15 conspicuous, translucent, silvery setae outside spiracle, extending towards posterior margin but not towards hind coxa; gaster dark brown, with a distinct coppery purple sheen, dorsal anterior part of Gt1 with a distinct dark blue, blue-green, brassy and coppery sheen, side of gaster with a mixed metallic green, brassy and coppery sheen; gonostylus dark brown.

Head (Fig. 1761) about 2.5X as wide as frontovertex, in profile about 1.7X as high as deep; occipital margin sharp, carinate; frontovertex with irregular, fairly deep, polygonal sculpture, in ocellar area of mesh size smaller than eye facet, subequal or conspicuously larger in front of anterior ocellus; ocelli forming an angle of about 105°; frontovertex with piliferous punctures shallow and indistinct, about 4 pairs of setae in front of anterior ocellus, a line of sparse setae along inner eye margin continued along lower eye margin on to gena; interantennal prominence with a few inconspicuous setae; temple and gena with moderately deep, longitudinally elongate, polygonally reticulate sculpture of relatively large mesh, area immediately below eye completely smooth; eye nearly reaching occipital margin, separated by about 0.5X diameter of facet, clothed with fairly inconspicuous setae, each clearly shorter than diameter of facet, separated from scrobal margin by very slightly less than diameter of ocellus, area between eye and scrobe partly with shallow, transversely elongate, polygonally reticulate sculpture; scrobes shallow, more or less ∩-shaped, meeting dorsally, very weakly margined dorsally and laterally; interantennal prominence shallow, dorsally rounded; antenna as in Fig. 1757; scape broadened and flattened, about 3.3X as long as broad, dorsal margin straight; all funicle segments at least a little longer than broad, becoming slightly broader distad; clava hardly enlarged, less than 1.1X as wide as F6, a little shorter than F4-F6 combined, outer suture not oblique, sensory area hardly enlarged, at apex only giving it a rounded or slightly pointed appearance; malar sulcus absent; clypeal margin with a very weak, convexly rounded median lobe; mandible with two acute teeth and a broad, truncate, upper tooth. Relative measurements: HW 76, HH 69, FV 31, POL 15, OOL 3, OCL 5, AOL 8.5, EL 44, EW 32, MS 26, SL 30, SW 9.

Thorax (Fig. 1758) in dorsal view with pronotum clearly visible behind head, its posterior margin concave, with polygonally reticulate sculpture that is generally similar to that on anterior part of frontovertex; mesoscutum with slightly irregular, polygonally reticulate sculpture that is similar to that on pronotum, of conspicuously smaller mesh posteriorly; axilla with transversely elongate, polygonally reticulate to imbricate-reticulate sculpture that is of similar depth to that on posterior part of mesoscutum; scutellum dorsally convex, mostly with similar sculpture to that on posterior part of mesoscutum, but clearly coarser, and much deeper, this continued on to side and almost to apex, but clearly shallower; visible part of mesoscutum about 1.8X as broad as long; scutellum nearly 1.1X as long as broad, apex not flange-like; fore wing with venation and distribution of setae as in Figs 1755, 1759; costal cell with a single line of setae dorsally in apical 0.5X or so; mid tibial spur about as long as basitarsus; propodeum (Fig. 1758) medially about 0.16X as long as scutellum, with some distinct, but irregular sculpture, side posteriorly sharp above hind coxa, but not carinate, with a small triangular tooth a little above half way towards spiracle. Relative measurements: FWL 88, FWW 35; HWL 60, HWW 15.

Gaster with hypopygium reaching about 0.6X to apex; syntergum about 0.8X as long as mid tibia, with apex acute; ovipositor clearly exserted, the exserted part about 0.5X as long as mid tibial spur or about 0.2X as long as mid tibia.

Paratype. Funicle with linear sensilla only on F2-F6; hypopygium Fig. 1760; ovipositor Fig. 1756. Relative measurements: OL 131, GL 43.5 [MT 78, MTS 27].

Variation. Females vary in overall length from 1.71-1.87mm, F1 may have linear sensilla, otherwise very little variation noted in material available.

Male. Unknown.

DISTRIBUTION. Costa Rica.

HOSTS. Unknown.

MATERIAL EXAMINED.

Type material. Holotype E: COSTA RICA, San José, 26km N San Isidro , 9°30’N 83°43’W, vi-vii.1992 (P. Hanson) . Paratypes: COSTA RICA, 2E, Guanacaste, ZP Miravalles, Las Hornillas , LN 299400 409200, 900m, #79677, 21.vii.2004, (J. Azofeifa); 2E , San José, Zurqui de Moravia , 1600m, v.1992 and iv.1995 (P. Hanson); 1E, San José, San Pedro, UCR Campus, 1200m, 28.ii.1997 (C. Hansson); 1E, San José, 26km N San Isidro, 9°30’N 83°43’W, 2100m, i-ii.1991 (P. Hanson, C. Godoy); 1E, San José, 26km N San Isidro, 9°30’N 83°43’W, 2100m, xi1992- i.1993 (P. Hanson); 1E, San José, Cerro (de la) Muerte, 19km S, 3km W Empalme, 2600m, iv-vii.1992 (P. Hanson, C. Godoy) . Holotype in MZUCR, paratypes in MZUCR and NHMUK .

COMMENTS. See comments under carme (p. 642).