Cleome amblyocarpa Barratte & Murbeck

Roalson, Eric H., 2021, A revised synonymy, typification, and key to species of Cleome sensu stricto (Cleomaceae), Phytotaxa 496 (1), pp. 54-68 : 58

2. Cleome amblyocarpa Barratte & Murbeck in Murbeck (1905: 25).

Lectotype (designated here): Tunisie: Maroc: Oasis d’Akka (plusieurs indiv. Recoltes par le rabbin Mardochee en mars 1872 et en 1873 [Hb. Coss.] ( P barcode P05328491 !; isolectotypes: LE barcode LE00013126 [photo!], P barcodes P05328488 !, P05328490 ! & P05328493 !). Additional syntypes: Algerie: Prov. d’Alger: Guerrara, autour de Beni Isguen dans l’Oued Mzab, 11 janvier 1857 [Hb. Coss.] ( P barcode P05328492 !); Algerie: Prov. de Const. : Lit d’un oued desscche pres de Beni Mora entre Biskra et Hammam Salahin, 15 avril 1903, S. Murbeck s.n. ( LD barcode LD1221985 [photo!]); Tunisie: Dj. Hattig, 17 mai, Miss. Bot. 1884 [Hb. Mus. Par.] (not found).

Homotypic synonym: Cleome arabica Linnaeus subsp. amblyocarpa (Barratte & Murbeck) Ozenda (1958: 247) .

Heterotypic synonyms: Siliquaria glandulosa Forsskal (1775: 78) . Type: Egypt: Cairo (‘ad Birket el hádgi prope Káhiram’), 1762, P. Forsskal 640 (lectotype, designated here: C barcode C10003078 [photo!]; isolectotype: C barcode C10003079 [photo!]). Cleome siliquaria Brown (in Salt 1814: 65). Cleome africana Botschantzev (1964: 130) . Cleome amblyocarpa Barratte & Murbeck var. glandulosa (Forrskal) Botschantzev (1968: 237) .— Cleome hexandra Poiret ex Steudel (1821: 206) , pro. syn.— Cleome arabica Linnaeus var. lutea Parsa (1947: 17) . Type: Bampour, 600 m. 6.5.39, Parsa 343 (holotype: K barcode K000643949!). Cleome daryoushiana Parsa (1951: 909) .

Note 1: Barratte and Murbeck (1905) clearly designate four collections on which this name is based (syntypes). Collections of three of these syntypes have been located, but the other has yet to be discovered. Despite this, six sheets are known that are syntype material. Botschantzev (1968) designated as the lectotype the Oasis d’Akka material with the lectotype at P and an isolectotype at LE. However, there are four sheets of the Oasis d’Akka collection at P, and so Botschantzev’s lectotypification is insufficient to clarify the type specimen. For this reason, I have chosen one of these ( P barcode P05328491 !) as the best representative of the material available and the lectotype .

Note 2: As described by Botschantzev (1964), Cleome siliquaria Brown cannot be used for this species because when the combination was made in Cleome , the specific epithet from the original publication was not kept, and there was not a prior published C. glandulosa so it cannot be considered a replacement name. Later, de Candolle (1824) applied the Ruiz and Pavon name Cleome glandulosa for a different species, but this was after the name change by Brown (in Salt 1814).

A second sheet of the type collection has been added to the Forsskal Herbarium at C. It is numbered 1735, but it seems clear this is the same collection as Forsskal 640, here designated the lectotype, as they are both collected in 1762 from “ad Birket el hádgi prope Káhiram” and it is here considered an isolectotype .

Geographic distribution: North Africa from Morocco east to Egypt, south to Sudan and Ethiopia, Arabian Peninsula, and throughout most of southwest Asia as far east as Iran.