Dichotomius mamillatus ( Felsche, 1901 )

Rosa, Cecilia Lozano De La, Cupello, Mario & Vaz-De-Mello, Fernando Z., 2024, The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution, European Journal of Taxonomy 959 (1), pp. 1-272 : 102-104

Dichotomius mamillatus ( Felsche, 1901)

Fig. 41A

Pinotus mamillatus Felsche, 1901: 143 (original description). Type locality: Ecuador. Name-bearing type: lectotype (SMTD), designated by Rossini & Vaz-de-Mello (2020), examined by FZVM.

Pinotus calcaratus Arrow, 1913: 457 (original description). Type locality: Brazil: Rondônia: Madeira River, “Madeira-Mamoré Railway Company Camp 41” (“Matto Grosso”, “Madeira-Mamore / R.R.Co.Camp 41”). Name-bearing type: holotype (BMNH) ( Rossini & Vaz-de-Mello 2020), examined by FZVM.

Pinotus mamillatus – Gillet 1911b: 61 (catalogue). — Balthasar 1941: 349 (checklist); 1951: 334 (checklist). — Blackwelder 1944: 207 (checklist). — Sarmiento-Garcés & Amat-García 2009: 291 (cited). — Rossini & Vaz-de-Mello 2020: 60 (cited).

Pinotus calcaratus – Luederwaldt 1929: 97, 102–103 (key, diagnosis); 1936: 213 (cited). — Rossini & Vaz-de-Mello 2020: 60 (cited as synonym of Dichotomius mamillatus ).

Dichotomius (Dichotomius) mamillatus – Vulcano & Pereira 1967: 585 (key, synopsis). — Luederwaldt 1929: 53 (key). — Amézquita et al. 1999: 119–120, 123 (list for Orinoquía, Colombia). — Escobar 2000: 208 (checklist for Colombia). — Medina & Lopera-Toro 2000: 309 (key for the Amazon). — Vaz-de-Mello 2000: 193 (checklist for Colombia). — Medina et al. 2001: 138 (checklist for Colombia). — Pulido-Herrera et al. 2003: 54 (list for Caqueta, Colombia); 2007: 307 (list for Andean Region Colombia). — Celi et al. 2004: 44 (list for Morona Santiago, Ecuador). — Noriega-Alvarado 2004: 39 (list for La Macarena, Colombia). — Noriega & Botero-Trujillo 2008: 451 (comments). — Noriega et al. 2008b: 79 (ecology). — Carpio et al. 2009: 469 (ecology). — Medina & Pulido-Herrera 2009: 60 (list for Orinoquia, Colombia). — Sarmiento-Garcés & Amat-García 2009: 287, 291, 295 (diagnosis, key). — Larsen 2011: 98, 102 (list for Suriname); 2013: 98 (list for Suriname). — Vaz-de-Mello et al. 2011b: 87 (list). — Krajcik 2012: 91 (checklist). — Noriega & Navarrete-Heredia 2013: 191 (comments). — Ratcliffe 2013: 493, 504, 510–511 (fig. 48, cited as Dichotomius nr. mamillatus ). — Martínez et al. 2014: 17 –18 (comments). — Silva et al. 2014: 349 (ecology); 2015: 611 (list for Mato Grosso state, Brazil); 2016: 554 (ecology, biogeography). — Ratcliffe et al. 2015: 196 (checklist for Peru). — Chamorro et al. 2018: 89 (key for Ecuador); 2019: 114–115 (catalogue). — Giraldo et al. 2018: 61 (guide). — Santos et al. 2018: 46 (list for Acre, Brazil). — Rossini & Vaz-de-Mello 2020: 59–61 (key, distribution, diagnosis, redescription).

Dichotomius (Dichotomius) calcaratus – Vulcano et al. 1976: 517 (new combination). — Vaz-de-Mello 2000: 193 (checklist for Brazil). — Rossini & Vaz-de-Mello 2020: 60 (as synonym of Dichotomius mamillatus ).

Dichotomius (Dichotomius) mamillatus – Vaz-de-Mello 2000: 193 (checklist for Colombia). — Vaz-de-Mello et al. 2011b: 87 (list). — Chamorro et al. 2018: 89 (key for Ecuador); 2019: 114–115 (catalogue). — Rossini & Vaz-de-Mello 2020: 59–61 (key, distribution, diagnosis, redescription).

Distribution

Colombia, Venezuela, Guyana, Suriname, French Guiana, Brazil, Ecuador, Peru, and Bolivia.

Subregions of Venezuela

Peneplain of the Casiquiare River –Upper Orinoco, System of hills and low piedmont mountains of the Guiana Shield, and Guiana Shield.

Literature record

Rossini & Vaz-de-Mello 2020: 60 ( Venezuela: Amazonas and Bolívar).

Figures

Fig. 41. Species distribution.A. Dichotomius mamillatus (Felsche, 1901).B. Dichotomius nisus (Olivier, 1789). Green diamond = CEMT collection data; black circle = literature data.