Dichotomius (Dichotomius) mamillatus (Felsche, 1901)

Chamorro, William, Marin-Armijos, Diego, senjo, Angelico & Vaz-De-Mello, Fernando Z., 2019, Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records, ZooKeys 826, pp. 1-343 : 110-112

Dichotomius (Dichotomius) mamillatus (Felsche, 1901) Plate 25B

Pinotus mamillatus Felsche, 1901: 143 (original description. Type locality: Ecuador).

Pinotus mamillatus : Gillet 1911a: 61 (complete list of species); Luederwaldt 1929: 53 (characters in key); Balthasar 1941: 349 (cited for Peru); Blackwelder 1944: 207 (list of species from Latin America); Balthasar 1951: 334 (cited for Peru).

Dichotomius mamillatus : Vulcano and Pereira 1967: 585 (characters in key); Vaz-de-Mello 2000: 193 (cited for Brazil); Medina et al. 2001: 138 (cited for Colombia); Sarmiento-Garcés and Amat-García 2009: 291 (diagnosis), 295 (characters in key); Carvajal et al. 2011: 320-321 (cited for Ecuador); Krajcik 2012: 91 (complete list of species); Sarmiento-Garcés and Amat-García 2014: 53 (characters in key), 73 (diagnosis); Ratcliffe et al. 2015: 196 (cited for Peru).

Dichotomius (Dichotomius) mamillatus : Chamorro et al. 2018: 94 (cited for Ecuador).

Types specimens.

Pinotus mamillatus Felsche, 1901. One syntype (♂) examined deposited at the SMTD. Lectotype to be designated in a future work on this species group.

Syntype (♂): "Ecuador / Baron [p]", "Typus [p, red label, black margin]", "Coll. C. Felsche / Kauf 20, 1918 [p, green label, black margin]", "mamillatus / Felsche / Ecuador [hw, purple label]", "Staatl. Museum für / Tierkunde D esden [p]".

Distribution.

Brazil, Colombia, Ecuador, and Peru.

Records examined.

MORONA SANTIAGO: Cumpi, Cordillera del Kutukú (1 specimen MUTPL); road Mendez-Paute, 1250 m (2 specimens CEMT; 5 specimens MQCAZ); NAPO: Nor Oeste de Puerto Napo, 470 m, Pungarayacu (1 specimen MQCAZ); Tena (3 specimens CEMT). ORELLANA: Bloque 31 Parque Nacional Yasuní (5 specimens MECN); Dayuma Campo Hormiguero, plataforma Hormiguero, 320 m (1 specimen MUTPL); Dayuma Campo Palanda-Yuca Sur plataforma Yuca 13, 255 m (1 specimen MGO-UC); Dayuma plataforma Ungurahua, 300 m (1 specimen MUTPL); El Dorado plataforma Guarango, 300 m (1 specimen MUTPL); Estación Científica Yasuní Puce, 250 m (19 specimens MQCAZ); Estación de Biodiversidad Tiputini, 285 m, Parque Nacional Yasuni (7 specimes MUTPL); Rodrigo Borja IAMOE (1 specimen CEMT; 9 specimens MQCAZ); San Sebastian del Coca, 345 m, Comuna Guataraco, Campo Pata (1 specimen MUTPL); San Sebastián del Coca, 345 m, Comuna Shamanal, Campo Palo Azul (1 specimen MUTPL). PASTAZA: Bosque Protector Oglán Alto, 660 m (1 specimen MUTPL); Chuyayacu, 200 m, Oleoducto km 25 (1 specimen MUTPL); Nuevo San Jose del Curaray, 245 m (2 specimens MUTPL); Pandanuque, 420 m (1 specimen MUTPL); San Virgilio (1 specimen MUTPL); Villano (3 specimens MECN). SUCUMBÍOS: Limoncocha, 250 m (2 speciemens CEMT); 6 km de Dureno, Precooperativa Los Vergeles, 290 m (4 specimens MGO-UC); Aucayacu Río El Eno, 275 m, 16 km de Lago Agrio (2 specimens MGO-UC); Nueva Loja plataforma Iguana, 310 m (1 specimen MUTPL); Pacayacu, 265 m, Campo Libertador, Tapi (1 specimen MUTPL); Tarapoa Campo Marian, plataforma Fanny 5, 260 m (1 specimen MUTPL); Tarapoa, Nuevo Manabí, 270 m (1 specimen MUTPL); Zabalo, 520 m (1 specimen CEMT). ZAMORA CHINCHIPE: Tundayme campamento, 800 m (2 specimens MUTPL); Tundayme campamento Mirador, San Marcos, 900 m (1 specimen MUTPL); Tundayme campamento Mirador, Las Maravillas, 1060 m (1 specimen COSEC); Zurmi, Comunidad La Wants, 1010 m (2 specimens MEPN; 1 specimen MUTPL); Zurmi Las Orquideas Río Nangaritza, 870 m (1 specimen MUTPL). UNDETERMINED PROVINCE: without specific locality (1 specimen SMTD).

Temporal data.

Collected every month of the year.

Remarks.

Inhabits the lowland evergreen forests, varzea forests and the foothill evergreen forests of the Amazon region from 200-1060 m a.s.l. Collected with flight interception traps and pitfall traps baited with carrion and human feces.