Thylogale eugenii decres Troughton, 1941

Parnaby, Harry E., Ingleby, Sandy & Divljan, Anja, 2017, Type Specimens of Non-fossil Mammals in the Australian Museum, Sydney, Records of the Australian Museum 69 (5), pp. 277-420 : 340

Thylogale eugenii decres Troughton, 1941

Furred Animals of Australia, 1st edition, p. 194. (20 November 1941).

Common name. Tammar Wallaby.

Current name. Notamacropus eugenii eugenii ( Desmarest, 1817a) , following Eldridge & Coulson (2015). Jackson & Groves (2015) elevated Notamacropus to generic rank and highlighted the need for further taxonomic resolution of the status of names included within N. eugenii . Eldridge et al. (2017) demonstrated that the Kangaroo Island population was genetically distinct, but its taxonomic status remains unresolved as they were unable to include definitive samples from extinct South Australian mainland populations.

Syntypes. (5, by subsequent determination): M.2821, male, skull, study skin, collected 13 February 1920 by E. Le G. Troughton, Deep Creek, 20 miles from Kingscote, Kangaroo Island , South Australia, registered 31 July 1920 ; M.2820, male, subadult, skull, study skin, collected 8 February 1920, other details as per M.2821; M.5332, female, adult, skull, study skin, (ex Taronga Zoo, Sydney), from Kangaroo Island , registered 16 March 1933 ; S.1808 and S.1809, both male skulls, collection dates not given, Kangaroo Island , registered 18 June 1928 collected by A. S. Le Souef.

Condition. M.2821: Cranium missing last erupted upper right molar and right alisphenoid bone, hole in left maxilla in the orbit area; right dentary missing coronoid and condylar processes. Study skin complete, in good condition. M.2820: Cranium and dentaries complete, in good condition; study skin missing tail tip and scrotum. M.5332: Cranium and dentaries complete. Study skin in good condition, slither of skin missing from the right ear. S.1808: Cranium missing left 3rd upper incisor; both dentaries complete. S.1809: Cranium and both dentaries complete.

Cranial measurements (mm). M.2821: GL, 90.64; ConL, 84.87; BasL, 79.59; NasL, 31.12; NasB, 14.85; DIL, 17.82; APV, 3.95; PPV, —; PAL, —; UPM (alv.), 4.42; UMR (alv.), 16.76 (M 1 to M 3); ZB, 49.07; POC, 17.10; MB, 37.19; DL (condyl.), 59.58 (left dentary); LPM (alv.), 2.82; LMR (alv.), 16.57 (M 1 to M 3). M.5332: GL, 97.83; ConL, 92.45; BasL, 87.86; NasL, 31.46; NasB, 17.07; DIL, 23.89; APV, 4.84; PPV, 12.69; PAL, 55.59; UPM (alv.), 4.13; UMR (alv.), 23.12 (M 1 to M 4); ZB, 51.20; POC, 16.22; MB, 40.71; DL (condyl.), 70.23; LPM (alv.), 3.61; LMR (alv.), 22.41 (M 1 to M 4). S.1808: GL, 101.79; ConL, 96.42; BasL, 90.36; NasL, 33.68; NasB, 18.63; DIL, 22.85; APV, 4.09; PPV, 11.86; PAL, 57.02; UPM (alv.), 4.64; UMR (alv.), 17.61 (M 1 to M 3); ZB, 54.24; POC, 16.34; MB, 40.21; DL (condyl.), 71.51; LPM (alv.), 3.88; LMR (alv.), 16.55 (M 1 to M 3). S.1809: GL, 101.95; ConL, 96.88; BasL, 92.54; NasL, 36.62* (tips worn); NasB, 17.51; DIL, 21.74; APV, 5.45; PPV, 10.51; PAL, 56.95* (posterior margin slightly eroded); UPM (alv.), 5.17; UMR (alv.), 17.08 (M 1 to M 3); ZB, 53.16; POC, 18.20; MB, 42.06; DL (condyl.), 71.05; LPM (alv.), 4.12; LMR (alv.), 16.60 (M 1 to M 3). [* = estimate].

Type localities. Deep Creek, 20 miles from Kingscote, Kangaroo Island , South Australia. Other syntypes are from “Kangaroo Island” .

Comments. Troughton (1941) stated that the subspecies was based on “specimens in the Australian Museum, including a male and female collected by the author in 1920” but does not cite registration or field numbers nor indicate the number of specimens in the type series. Calaby & Richardson (1988) listed registration numbers of the five specimens listed above, which they recognize as syntypes. Troughton did not nominate a holotype in his description, but the specimen tag of M.2821 is labelled holotype, M.5332 as allotype and M.2820 as paratype, all in Troughton’s characteristic handwriting; they are listed here as syntypes because Troughton’s attributions remain unpublished. Specimen index cards for S.1808–9 are not marked as paratypes but they are recognized as syntypes because they were available to Troughton in 1941. Publication date drawn from Whitley (1975).