Dichotomius (Selenocopris) fortepunctatus ( Luederwaldt, 1923 )

Chamorro, William, Lopera-Toro, Alejandro & Rossini, Michele, 2021, A new species and distribution records of Dichotomius Hope, 1838 (Coleoptera Scarabaeidae: Scarabaeinae) in Colombia, Zootaxa 4942 (2), pp. 193-206 : 203

Dichotomius (Selenocopris) fortepunctatus ( Luederwaldt, 1923)

( Figs. 4E , 5 )

Pinotus fortepunctatus Luederwaldt, 1923: 4 (original description). Type locality: Bucay, west-Ecuador 300 m.

Pinotus fortepunctatus: Blackwelder 1944: 207 (checklist); Pereira 1953: 387 (synonym of Dichotomius globulus ( Felsche, 1901) , comment).

Dichotomius (Dichotomius) fortepunctatus: Chamorro et al. 2019a: 127–129 (designation of lectotype, revalidated name, records for Ecuador).

Type series. The lectotype (male) is deposited at the MZUSP ( Chamorro et al. 2019a: 127).

Records examined. COLOMBIA: NARIÑO: Tumaco, La Espriella 50 m, human feces, F. Escobar. VIII.1994 (1 specimen IAvH) .

Literature records. ECUADOR: CARCHI: Tobar Donoso , 300 m. COTOPAXI: Guasaganda km 4, 500 m. ESMERALDAS: Colón del Onzole, Charco Vicente, Gualpi, El Pajonal, El Progreso, Salto del Bravo , GUAYAS: Bucay. IMBABURA: El Chontal, El Cauchero, 900 m. LOS RÍOS: 47 km S Santo Domingo, Río Palenque Station, 250 m, Río Palenque Station . PICHINCHA: Bosque Protector Milpe-Río Pachijal, 1200 m, El Encuentro, 620 m, San Miguel de los Bancos, La Florida, Guayabilla 520 m, Río Guayllabamba Manduriacus, Mindo, Pedro Vicente Maldonado 600 m, Llurimaguas 290 m, Río Guayllabamba Pedro Vicente Maldonado , Tortugo (450 m), Río Guayllabamba Pedro Vicente Maldonado ; San Roque (580 m), Río Guayllabamba Pedro Vicente Maldonado . SANTO DOMINGO DE LOS TSÁCHILAS: Hacienda Pupusa, Río Silanche, Santo Domingo , Puerto Limón , 395 m. (see distribution data in Luederwaldt 1923: 4; Chamorro et al. 2019: 128) .

Remarks. Medina et al. (2001) reported 36 Dichotomius species from Colombia. Subsequently, Sarmiento-Garcés &Amat-García (2014), in their monograph on Colombian Dichotomius , provided some updates on the genus and reduced the species number to 32. Recently, Nunez & Vaz-de-Mello (2019) recorded Dichotomius (Cephagonus) fonsecae ( Luederwaldt, 1926) from the Chiribiquete National Park, in southern Colombia. This species was cited by Sarmiento-Garcés & Amat-García (2014) as “ D. (Selenocopris) sp.”, which until then was only known from foothill evergreen forests of the Amazon region of Ecuador. Finally, Montoya & Vaz-de-Mello (2019) described the male of Dichotomous foveicollis (Kirsch, 1871) , whose type locality is Bogotá.

Dichotomius (Selenocopris) fortepunctatus ( Luederwaldt, 1923) is recorded for the first time from Colombia. With this new distributional record, its geographical range now includes the biogeographical provinces of western Ecuador and Cauca (sensu Morrone 2014) ( Fig. 5 ).

Figures

FIGURE 4. Dichotomius species. A, Dichotomius podalirius, aedeagus; B, Dichotomius podalirius, endophallites; C, Dichotomius prietoi (dorsal view); D, Dichotomius fortepunctatus (dorsal view).

FIGURE5. Distributionof Dichotomius (Dichotomius) quadrilobatusnewspecies(greencircle) andDichotomius(Selenocopris) fortepunctatus (Luederwaldt, 1923) (blue square). (?) Doubtful data.