Antillocladius Saether

Mendes, Humberto F., Andersen, Trond & Hagenlund, Linn K., 2011, New species and records of Antillocladius Saether and Litocladius Mendes, Andersen et Saether from Brazil and Costa Rica (Chironomidae: Orthocladiinae), Zootaxa 2915, pp. 39-51 : 40-41

Key to the males of Antillocladius Saether

1. Squama bare. Mexico.............................................. A. zempoalensis Mendes, Andersen et Saether

- Squama with at least one seta............................................................................ 2

2. Anal point nearly parallel-sided, with numerous, weak, partly anteriorly directed setae; inferior volsella either pointed triangu- lar or consisting of a long apically simple or bifid lobe; wing bare............................................... 3

- Anal point tapering, with numerous, usually strong and posterolaterally directed setae; inferior volsella either with a dorsal anterior triangular to digitiform part and a more rounded ventral lobe or circular with or without additional rounded posterior extension, and conspicuously set off; wing usually with apical setae.............................................. 9

3. Inferior volsella pointed triangular; costal extension> 1.5 times RM............................................ 4

- Inferior volsella consisting of a long posteromedially directed simple or apically bifid lobe; costal extension <1.5 times RM..................................................................................................... 6

4. Inferior volsella triangular, leaf-like. Brazil..................................... A. folius Mendes, Andersen et Saether

- Inferior volsella sharply pointed or with rounded apex........................................................ 5

5. Inferior volsella small, sharply pointed. Brazil.................................. A. biota Mendes, Andersen et Saether

- Inferior volsella large, with rounded apex. Brazil.................................... A. axitiosus Mendes et Andersen

6. Inferior volsella apically bifid............................................................................ 7

- Inferior volsella simple.................................................................................. 8

7. Virga absent. Brazil................................................... A. sooretama Mendes, Andersen et Saether

- Virga present. Brazil........................................................... A. atalaia Mendes et Andersen

8. Inferior volsella uniformly colored, with sharply pointed apex. Brazil, Venezuela.... A. ubatuba Mendes, Andersen et Saether

- Inferior volsella with a distinctly darker oral part bearing strong setae, with rounded apex. Brazil................................................................................................... A. ultimus Mendes et Andersen

9. Virga longer than half the length of phallapodeme........................................................... 10

- Virga absent or shorter than half the length of phallapodeme................................................... 17

10. Wing membrane bare; costal extension about 2 times the length of RM. Brazil....................... A. anandae sp. n.

- Wing membrane with at least a few seta in cell r4+5, costal extension <1.5 times RM............................... 11

11. Anal point with numerous weak setae; inferior volsella small. Venezuela, Ecuador................................................................................................. A. venequatoriensis Mendes, Andersen et Saether

- Anal point with strong, stiff setae; inferior volsella well developed.............................................. 12

12. Wing membrane with <10 setae apically in cell r4+5; AR 1.4–1.7............................................... 13

- Wing membrane with> 20 setae apically in cell r4+5, setae also present apically in cell m1+2 and sometimes in cell m3+4; AR 0.9– 1.6................................................................................................ 14

13. Virga nearly as long as phallapodeme. Brazil.................................. A. musci Mendes, Andersen et Saether

- Virga less than half the length of phallapodeme. China, Thailand............................ A. zhengi Wang et Saether

14. Virga about two-thirds the length of phallapodeme.......................................................... 15

– Virga half the length of phallapodeme.................................................................... 16

15. Squama with 5–6 setae; cell m3+4 with several setae. Ecuador, Mexico, Nicaragua, U.S.A............ A. pluspilalus Saether

- Squama with 9–13 setae; cell m3+4 bare. Japan......................................... A. yakyijeus (Sasa et Suzuki)

16. Cell m3+4 bare; AR 1.22–1.56. China, Russia....................................... A. scalpellatus Wang et Saether

- Cell m3+4 with setae; AR about 1.0 (Holotype). India............................ A. subnubilus (Sinharay et Chaudhuri)

17. Virga consisting of 2 or 6 very short spines................................................................ 18

- Virga absent......................................................................................... 23

18. Virga consisting of 6 spines; inferior volsella with digitiform dorsal anterior lobe covering rounded posteroventral lobe. Mex- ico............................................................. A. calakmulensis Mendes, Andersen et Saether

- Virga consisting of 2 spines............................................................................. 19

19. Inferior volsella apically split in 2 semicircular lobes. Mexico................ A. herradurus Mendes, Andersen et Saether

- Inferior volsella not divided into lobes.................................................................... 20

20. Wing with protruding anal lobe.......................................................................... 21

- Anal lobe weak or absent.............................................................................. 22

21. Inferior volsella with median, oblique fold; anterior acrostichals simple, posterior scalpellate. Brazil................................................................................................ A. plicatus Mendes et Andersen

- Inferior volsella simple; all acrostichals scalpellate. Brazil......................................... A. itatiaia sp. n.

22. Costa distinctly extended; inferior volsella perpendicular to gonocoxite; setae on anal point weak. Brazil............................................................................................ A. brazuca Mendes et Andersen

- Costa not extended; inferior volsella inclined in relation to gonocoxite; setae on anal point strong. Japan........................................................................................... A. tokarameneus (Sasa et Suzuki)

23. Inferior volsella well set off and circular or weak and adpressed to gonocoxite; wing membrane usually with setae....... 24

- Inferior volsella well set off, parallel-sided or circular, with additional posterior semicircular extension; wing membrane bare.................................................................................................... 26

24. Inferior volsella with dorsal ridge. Brazil......................................... A. gephyrus Mendes et Andersen

- Inferior volsella simple................................................................................ 25

25. Wing membrane with numerous apical setae, present also in cell m3+4. Ecuador... A. skartveiti Andersen et Contreras-Ramos

- Wing membrane with 0–3 setae apically in cell r4+5. Brazil, Costa Rica, Saint Vincent, Venezuela....... A. antecalvus Saether

26. Inferior volsella well set off, circular, with additional posterior semicircular extension; costal extension> 54 µm long. Brazil, Costa Rica, Mexico, Venezuela, U.S.A....................................................... A. arcuatus Saether

- Inferior volsella well set off, parallel-sided, with broadly rounded apex; costal extension <45 µm. Brazil.......................................................................................... A. campususp Mendes et Andersen