Saphenista Walsingham, 1914
publication ID |
https://doi.org/ 10.5281/zenodo.3676593 |
publication LSID |
lsid:zoobank.org:pub:68DF38B0-C69E-4394-8992-CD74633ECADA |
DOI |
https://doi.org/10.5281/zenodo.3681445 |
persistent identifier |
https://treatment.plazi.org/id/EB0C87B8-FF8F-4017-A8BC-B408FBDAFB95 |
treatment provided by |
Felipe |
scientific name |
Saphenista Walsingham, 1914 |
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Saphenista Walsingham, 1914 View in CoL
Type species. Conchylis lacteipalpis Walsingham, 1891 , by original designation (Walsingham 1914).
The precise circumscription of Saphenista View in CoL and its putative synonyms (see Brown 2005) has been somewhat elusive owing to the absence of an abdomen of the holotype of the type species of the genus (i.e., Conchylis lacteipalpis ). As currently defined, the genus is anchored by species that are superficially similar to its type species and assigned to the genus by Walsingham (1914). The most convincing morphological autapomorphy for Saphenista View in CoL is the conspicuous Y- or T-shaped median process of the transtilla, which easily separates males from those of all other Cochylina. Razowski (1994) identified the following as putative autapomorphies for the genus: a tooth-like process of the vinculum (highly variable and not always present); a sclerotized pocket at the base of the valva (usually present); an expanded apex of the median process of the transtilla (mentioned above, always present); and the presence of a subterminal prominence of the phallus (absent in many species). Also, males of some species of Saphenista View in CoL have secondary structures from the venter of the abdomen. For example, S. multistrigata Walsingham, 1914 View in CoL has a small bilobed process on abdominal segment 7 from which a pair of laterally directed hairpencils originate. Similar structures are reported and illustrated by Razowski and Becker (1983: fig. 3, 21) in S. nauphraga Razowski and Becker, 1983 View in CoL and S. ficta Razowski and Becker, 1983 View in CoL .
Based on morphological features, primarily the dorso-posteriorly directed socii, Razowski (1985, 1994) concluded that Saphenista View in CoL is closely related to Phalonidia View in CoL , and a recent phylogenetic analysis of the subtribe ( Brown et al. 2019) provides some support for this conclusion, although in the latter study Saphenista View in CoL (+ Amallectis Meyrick, 1917 View in CoL ) was recognized as one of six major lineages in the subtribe Cochylina, sister to the Phalonidia View in CoL Group + the Cochylis View in CoL Group (sensu Brown et al. 2019).
Adults of Saphenista View in CoL are highly variable in maculation and size (i.e., forewing length). Males lack a forewing costal fold, and most (but not all) species have a hindwing costal roll (hair pencil), the last feature shared with most “advanced” Cochylina ( Brown et al. 2019) . In contrast, females of many Saphenista View in CoL species retain the plesiomorphic condition of three spines in the frenulum.
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