Trichoptera,

Johanson, Kjell Arne & Oláh, János, 2012, Revision of the Fijian Chimarra (Trichoptera, Philopotamidae) with description of 24 new species, Zootaxa 3354, pp. 1-58: 6-7

publication ID

http://doi.org/ 10.5281/zenodo.210736

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http://treatment.plazi.org/id/9F3E87DD-562B-FFD0-E89A-FF5AFA17F95D

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Plazi

scientific name

Trichoptera
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Key to Fijian Trichoptera  families (adults) and Philopotamidae  species (males)

1 Forewings shorter than 4.0 mm, narrow, pointed apically (figs. 6–10 in Marshall (1979 )); antennae shorter than forewings; tho- rax with mesoscutellum pointed posteriorly (fig. 83 in Marshall (1979 ))................................................................................................. Hydroptilidae  (1 species: Oxyethira fijiensis Kelley, 1989  ).

- Forewings longer than 4.0 mm, broad, rounded apically ( Fig. 5View FIGURE 5); antennae as long as forewings or longer; thorax with mesoscutellum rounded or nearly straight posteriorly (figs. 53–54 in Marshall (1979 ))............................... 2

2 (1 ’) Ocelli present........................................................................................ 3

- Ocelli absent......................................................................................... 4

3 (2) Maxillary palps each with segment 2 swollen mesally and with apical segment flexible, distinctly longer than previous seg- ment............................................................. Philopotamidae  ( Chimarra  , 27 species)... 8

- Maxillary palps each with segment 2 cylindrical and with apical segment not flexible, about as long as previous segment...................................................................... Hydrobiosidae  ( Apsilochorema  , 3 species).

4 (2 ’) Maxillary palps each with apical segment flexible, distinctly longer than previous segment........................... 5

- Maxillary palps each with apical segment not flexible, about as long as previous segment........................... 6

5 (4) Forelegs each with 2 spurs (preapical spur absent); thorax without mesonotal setal warts................................................................................................... Hydropsychidae  ( Abacaria  , 9 species).

- Forelegs each with 3 spurs (preapical spur present); thorax with pair of mesonotal setal warts......................................................................................... Polycentropodidae  ( Polyplectropus  , 8 species).

6 (4 ’) Antennae each with scape distinctly longer than head; thorax with pair of distinct mesonotal setal warts, setae never in longitu- dinal rows..................................................................... Goeridae  ( Goera  , 3 species).

- Antennae each with scape shorter or about as long as head; thorax without mesonotal setal warts, setae always in pair of longi- tudinal rows.......................................................................................... 7

7 (6 ’) Maxillary palps each 6 -segmented; forewings less than 3 x longer than broad... Calamoceratidae  ( Anisocentropus  , 5 species)

- Maxillary palps 5 -segmented; forewings more than 3 x longer than broad................................................................................ Leptoceridae  ( Oecetis  , 1 species; Triaenodes  , 3 species; Triplectides  , 1 species)

8 (3) Hind wings each with centrally located, large, pale hyaline spot ( Figs. 10–13View FIGURES 6 – 13)..................................... 9

- Hind wings without large pale, hyaline spots ( Figs. 6–7View FIGURES 6 – 13).................................................... 14

9 (8) In genitalia, gonopods short, length about as long as high in lateral view ( Fig. 50View FIGURES 50 – 54).............. C. schlingeri  , new species.

- In genitalia, gonopods more elongate, longer than high in lateral view (cf. Figs. 60View FIGURES 60 – 64, 65View FIGURES 65 – 69, 70View FIGURES 70 – 74)......................... 10

10 (9 ’) In genitalia, ventral branch of each gonopod more than 2 x longer than wide in lateral view ( Fig. 60View FIGURES 60 – 64) C. braueri  , new species.

- In genitalia, ventral branch of each gonopod not as long as wide in lateral view ( Figs. 45View FIGURES 45 – 49, 65View FIGURES 65 – 69)........................ 11

11 (10 ’) In genitalia, gonopods widest at mid-length in lateral view ( Fig. 70View FIGURES 70 – 74)..................... C. karoyanitensis  , new species.

- In genitalia, gonopods widest beyond mid-length in lateral view ( Figs. 45View FIGURES 45 – 49, 55View FIGURES 55 – 59, 65View FIGURES 65 – 69)................................. 12

12 (11 ’) In genitalia, dorsal branch of each gonopod forming ventrally pointing hook in lateral view ( Fig. 45View FIGURES 45 – 49) C. signata Banks, 1936  .

- In genitalia, dorsal branch of each gonopod forming posteriorly pointing triangle in lateral view ( Figs. 55View FIGURES 55 – 59, 65View FIGURES 65 – 69)......... 13

13 (12 ’)In genitalia, dorsal margin of each gonopod nearly straight and as darkly sclerotized as posterior margin ( Fig. 55View FIGURES 55 – 59)........................................................................................... C. nathani  , new species.

- In genitalia, dorsal margin of each gonopod strongly convex and more darkly sclerotized than posterior margin ( Fig. 65View FIGURES 65 – 69)...................................................................................... C. vitiensis  , new species.

14 (8 ’) Forewings each with large, pale hyaline spot............................................................... 15

- Forewings without large, pale hyaline spots............................................................... 16

15 (14) Forewings each with large, pale hyaline spot located immediately anterior of posterior wing margin ( Fig. 6View FIGURES 6 – 13); in genitalia, ter- gum X with prominent dorsal process on each lateral branch ( Fig. 35View FIGURES 35 – 39)....................... C. vanuensis  , new species.

- Forewings each with large, pale hyaline spot located centrally in wings ( Fig. 7View FIGURES 6 – 13); in genitalia, tergum X without dorsal pro- cesses ( Fig. 40View FIGURES 40 – 44)................................................................. C. macuatensis  , new species.

16 (14 ’)In genitalia, phallic apparatus without endothecal processes; phallotremal sclerite more than 2 x longer than width of narrowest part of phallotheca ( Figs. 79View FIGURES 75 – 79, 84View FIGURES 80 – 84, 89View FIGURES 85 – 89)..................................................................... 17

- In genitalia, phallic apparatus with 1 or 2 pairs of endothecal processes; phallotremal sclerite less than 2 x longer than width of narrowest part of phallotheca, or not discernable ( Figs. 139View FIGURES 139 – 144, 170View FIGURES 170 – 174)............................................... 19

17 (16) In genitalia, gonopods acute apically in lateral view ( Fig. 75View FIGURES 75 – 79); phallotremal sclerite more than 1 / 2 as long as phallotheca ( Fig. 79View FIGURES 75 – 79)............................................................................. C. tokotaai  , new species.

- In genitalia, gonopods rounded apically in lateral view ( Figs. 80View FIGURES 80 – 84, 85View FIGURES 85 – 89); phallotremal sclerite much less than 1 / 2 as long as phal- lotheca ( Figs. 84View FIGURES 80 – 84, 89View FIGURES 85 – 89).................................................................................. 18

18 (17 ’)In genitalia, segment IX rectangular and oblique, with ventral margin much shorter than height in lateral view ( Fig. 85View FIGURES 85 – 89); each posterior margin produced into large, triangular plate immediately below its cercus ( Fig. 85View FIGURES 85 – 89).. C. naitasirensis  , new species.

- In genitalia, segment IX hyperbolic, with ventral margin subequal to height in lateral view ( Fig. 80View FIGURES 80 – 84); each posterior margin without triangular plate below its cercus ( Fig. 80View FIGURES 80 – 84)............................................ C. vuda  , new species.

19 (16 ’)In genitalia, phallic apparatus with 2 pairs of endothecal processes ( Fig. 138View FIGURES 134 – 138)..................................... 20

- In genitalia, phallic apparatus with 1 pair of endothecal processes ( Fig. 169View FIGURES 165 – 169)..................................... 23

20 (19) In genitalia, each lateral branch of tergum X with dorsal, rounded process ( Figs. 105View FIGURES 105 – 110, 111View FIGURES 111 – 116).......................... 21

- In genitalia, each lateral branch of tergum X with dorsal, hook-shaped process ( Figs. 134View FIGURES 134 – 138, 139View FIGURES 139 – 144)...................... 22

21 (20) In genitalia, anteroventral portion of segment IX rectangular in lateral view ( Fig. 105View FIGURES 105 – 110); in tergum X, gap between dorsal and ventral branches nearly right-angled ( Fig. 105View FIGURES 105 – 110); tergum X with triangular process on lateral margin in lateral view ( Fig. 105View FIGURES 105 – 110); gonopods strongly expanded ventrally at base ( Fig. 105View FIGURES 105 – 110); in phallus, endothecal processes no longer than as minimum breadth of phallotheca in ventral view ( Fig. 110View FIGURES 105 – 110)................................................. C. manni Banks, 1924  .

- In genitalia, anteroventral portion of segment IX triangular in lateral view ( Fig. 111View FIGURES 111 – 116); in tergum X, gap between dorsal and ventral branches wide-angled, nearly 180 degrees ( Fig. 111View FIGURES 111 – 116); tergum X without triangular process on ventral margin in lateral view ( Fig. 111View FIGURES 111 – 116); gonopods only slightly expanded ventrally at base ( Fig. 111View FIGURES 111 – 116); in phallus, endothecal processes longer than minimum breadth of phallotheca in ventral view ( Fig. 116View FIGURES 111 – 116)............................... C. cakaudrovensis  , new species.

22 (20 ’) In genitalia, each dorsal branch of tergum X with darkly sclerotized apicodorsal hook ( Fig. 134View FIGURES 134 – 138); each ventral branch of tergum X reaching as far posterior as each dorsal branch ( Fig. 134View FIGURES 134 – 138); phallic apparatus without spicules ( Figs. 137, 138View FIGURES 134 – 138)............................................................................................ C. lavensis  , new species.

- In genitalia, each dorsal branch of tergum X with weakly sclerotized subapicodorsal process ( Fig. 139View FIGURES 139 – 144); each ventral branch of tergum X exceeding posteriorly each dorsal branch of tergum X ( Fig. 139View FIGURES 139 – 144); phallic apparatus with group of spicules in addition to endothecal processes ( Figs. 143, 144View FIGURES 139 – 144)................................................ C. devoensis  , new species.

23 (19 ') In genitalia, tergum X with each lateral branch uniramus ( Figs. 90View FIGURES 90 – 94, 124View FIGURES 124 – 128, 155View FIGURES 155 – 159)..................................... 24

- In genitalia, tergum X with each lateral branch biramus ( Figs. 95View FIGURES 95 – 99, 117View FIGURES 117 – 123, 145View FIGURES 145 – 149)..................................... 29

24 (23) In genitalia, gonopods clearly reaching beyond tergum X posteriorly ( Figs. 100View FIGURES 100 – 104, 155View FIGURES 155 – 159)........................... ... 25

- In genitalia, gonopods reaching as far out as tergum X or shorter ( Figs. 90View FIGURES 90 – 94, 150View FIGURES 150 – 154, 164View FIGURES 160 – 164).............................. 26

25 (24) In genitalia, segment IX long, with narrow anteroventral plates and straight posterior margin in lateral view ( Fig. 100View FIGURES 100 – 104); each gonopod uniramus ( Fig. 100View FIGURES 100 – 104)....................................................... C. abacensis  , new species.

- In genitalia, segment IX short, with broad anteroventral plates and convex posterior margin in lateral view ( Fig. 155View FIGURES 155 – 159); each gonopod with slender dorsal branch and rounded ventral branch ( Fig. 155View FIGURES 155 – 159).................. C. psychodida  , new species.

26 (24 ’) In genitalia, gonopods about as long as broad in lateral view ( Fig. 90View FIGURES 90 – 94)....................... C. veisarensis  , new species.

- In genitalia, gonopods much shorter than broad in lateral view ( Fig. 124View FIGURES 124 – 128, 165View FIGURES 165 – 169, 170View FIGURES 170 – 174)................................ 27

27 (26 ') In genitalia, distal part of each gonopod almost straight ( Figs. 124View FIGURES 124 – 128, 170View FIGURES 170 – 174)........................................ 33

- In genitalia, distal part of each gonopod curved posterad in lateral view ( Figs. 150View FIGURES 150 – 154, 165View FIGURES 165 – 169)............................ 28

28 (27 ’) In genitalia, segment IX with slightly concave to straight posterior margin ( Fig. 150View FIGURES 150 – 154); tergum X short, not reaching beyond gonopods in lateral view ( Fig. 150View FIGURES 150 – 154); each lateral branch of tergum X with lateral subapical process ( Fig. 151View FIGURES 150 – 154); gonopods each with rounded ventral branch in lateral view ( Fig. 150View FIGURES 150 – 154)..................................... C. tipulida  , new species.

- In genitalia, segment IX with slightly convex posterior margin ( Fig. 165View FIGURES 165 – 169); tergum X long, reaching well beyond gonopods in lateral view ( Fig. 165View FIGURES 165 – 169); tergum X without lateral processes ( Fig. 166View FIGURES 165 – 169); gonopods each with nearly rectangular ventral branch in lateral view ( Fig. 165View FIGURES 165 – 169).............................................................. C. malickyi  , new species.

29 (23 ’) In genitalia, gonopods broad, nearly triangular in lateral view ( Fig. 117View FIGURES 117 – 123).................... C. cartwrighti  , new species.

- In genitalia, gonopods narrow, each with nearly parallel-sided dorsal branch ( Figs. 95View FIGURES 95 – 99, 129View FIGURES 129 – 133, 145View FIGURES 145 – 149, 160View FIGURES 160 – 164)................ 30

30 (29 ’) In genitalia, tergum X with ventral branches produced posterad beyond dorsal branches ( Figs. 95View FIGURES 95 – 99, 145View FIGURES 145 – 149)................ 31

- In genitalia, tergum X with ventral branches not produced posterad beyond dorsal branches ( Figs. 129View FIGURES 129 – 133, 160View FIGURES 160 – 164)............ 32

31 (30) In genitalia, apex of each dorsal and ventral branch of each gonopod parallel-sided and bent posterad C. fijiana  , new species.

- In genitalia, apex of dorsal branch of each gonopod straight; dorsal branch nearly parallel-sided, ventral branch irregular ( Fig. 145View FIGURES 145 – 149).......................................................................... C. helomyzida  , new species.

32 (30 ’) In genitalia, gonopods longer than segment IX in lateral view, slightly curved posterad along their length ( Fig. 129View FIGURES 129 – 133); distal 1 / 2 of tergum X narrower than distal 3 rd of gonopods in lateral view ( Fig. 129View FIGURES 129 – 133)................. C. kadavuensis  , new species.

- In genitalia, gonopods shorter than segment IX in lateral view, strongly curved into 1 / 2 -circle ( Fig. 160View FIGURES 160 – 164); tergum X broader than distal 3 rd of gonopods in lateral view ( Fig. 160View FIGURES 160 – 164)..................................... C. levuensis  , new species.

33 (27) In genitalia, posterolateral margins of segment IX each with posterior, setose lobe above its gonopod in lateral view ( Fig. 124View FIGURES 124 – 128); gonopods each narrowing at 1 / 2 -length into parallel-sided dorsal branch and with rounded ventral branch in lateral view ( Fig. 124View FIGURES 124 – 128); tergum X with convex dorsal margin in lateral view ( Fig. 124View FIGURES 124 – 128), divided into lateral processes in dorsal view ( Fig. 125View FIGURES 124 – 128).................................................................................. C. kimminsi  , new species.

- In genitalia, posterolateral margins of segment IX without posterior, setose lobes in lateral view ( Fig. 170View FIGURES 170 – 174); gonopods each with dorsal branch narrowing distally along its full length and with ventral branch short, triangular in lateral view ( Fig. 170View FIGURES 170 – 174); tergum X with stepped, concave dorsal margin in lateral view ( Fig. 170View FIGURES 170 – 174), without lateral processes in dorsal view ( Fig. 171View FIGURES 170 – 174)................................................................................. C. obscurella Banks, 1924  .