Phuphena Walker, 1858

Goldstein, Paul Z. & Zilli, Alberto, 2019, Thraumata, a new genus from South America with description of a new species from Peru (Lepidoptera, Noctuidae), ZooKeys 867, pp. 139-160 : 139

Phuphena Walker, 1858

Type species.

Phuphena fusipennis Walker, 1858 by monotypy.

Diagnosis.

Species of Phuphena diverge with respect to wing pattern, but most share a bounded medial area. Orbicular and reniform stigmata sometimes absent or indistinct; outer forewing margin smooth in the majority of species but crenulate in cilix . Hindwing upperside coloration uniform. Male genitalia of Phuphena distinct, the valvae uniquely reduced, with no evidence of clasping architecture, narrow for most of their length, and in most species swollen apically to form a knob-like cucullus, giving them a club-shaped appearance, with a corona comprising an unorganized arrangement of setae. Vinculum V-shaped, slightly truncate in some species, and extending below the base of the valves for a distance equal to or greater than the width at its broadest point. The phallus is asymmetrically sclerotized, often with a patch of scale-like cornuti, and the vesica without well-developed cornuti or diverticula, but with a corresponding ventro-distal strip of scale-like cornuti. Basal abdominal brushes present but not visibly subdivided as in Callopistria Hübner, [1821] (type species: Noctua juventina Stoll, 1782); a smaller pair on A8, and eversible saccular hair pencils completely absent. Female genitalia simple, the corpus bursae lemon-shaped, with no appendix or signa, the ductus seminalis attached apically, to the distal (anterior) end of the corpus. This distinguishes Phuphena from all Thraumata spp. and from Callopistria juventina , in which the ductus arises from the proximal (posterior) end of the corpus, but this character varies among other Callopistria species.