Glomeris

Golovatch, Sergei, Mauriès, Jean-Paul, Akkari, Nesrine, Stoev, Pavel & Geoffroy, Jean-Jacques, 2009, The millipede genus Glomeris Latreille, 1802 (Diplopoda, Glomerida, Glomeridae) in North Africa, ZooKeys 12 (12), pp. 47-86 : 83

Key to the known Glomeris species of North Africa

1 Tegument entirely pallid, rarely only ocelli dark. Cavernicoles ................... 2

– Head and terga distinctly pigmented, colour pattern evident, coloration mostly vivid. Epigean................................................................................. 3

2 Collum with a single transverse stria. Syncoxital horns of telopod clearly bifid ( Fig. 2D, E ). Libya..................................................... G. monostriata sp. n.

– Collum with the usual two transverse striae. Syncoxital horns of telopod surmounted with a setoid filament ( Fig. 1F ). Kabylia, Algeria........................... ............................................................................ G. troglokabyliana sp. n.

3 Colour pattern of terga 2-11 without evident, light, paramedian spots, but with alternating light and dark transverse bands (Figs 10, 12, 24) .............. 4

– Colour pattern of terga 2-(6)11 with 1+1 or 2+2 more or less evident light spots ( Figs 3 , 4 , 7 , 15-17 , 20-22, 26, 27) ................................................... 5

4 Coloration of caudal halves of terga distinctly marbled throughout (Figs 10, 12)........................................................................................... G. klugii

– Transverse bands on terga at most only slightly marbled in contact zones ( Fig. 24 ).......................................................................... G. mohamedanica

5 Unusually large yellowish to orange spots on tergum 6, following terga 7-11 uniformly blackish ( Figs 20-22 ) ...................................... G. carthaginiensis

– Spots on tergum 6 of same size as on adjacent terga ................................... 6

6 Median spots on tergum 2 much smaller than lateral ones ..... G. anisosticta

– Either both pairs of spots on tergum 2 comparable in size or lateral ones (if present) smaller than median ones ............................................................. 7

7 Coloration uniformly blackish, lighter spots (1+1 or 2+2) very vague, marbled (Figs 15-18)................................................................... G. sublimbata

– Background coloration brown to blackish, lighter spots more distinct........ 8

8 Tergum 2 with a single central light spot against a black-brown background. Pygidium uniformly black-brown. Morocco......................... G. brolemanni

– Tergum 2 with 2+2 light spots ................................................................... 9

9 Background coloration light to dark brown, drab, pattern as in Figs 26 , 27 .................................................................................................... G. punica

– Background coloration dark brown to blackish, pattern vivid .................. 10

10 Light axial stripe absent ( Figs 6-8 ).................................... G. flavomaculata

– Light axial stripe present ( Figs 3 , 4 ).................................. G. colorata sp. n.

Figures

Figure 1. Glomeris troglokabyliana sp. n., ♁ paratype; A, left part of head showing ocelli and Tömösváry’s organ, dorsal view; B, antenna, lateral view; C, thoracic shield and terga 3 & 4, lateral view; D, leg 17; E, leg 18; F, G, leg 19 (telopod), caudal and frontal views, respectively. – Scale bar: 0.3 mm (D-G); drawn not to scale (A-C).

Figure 2. Glomeris monostriata sp. n., ♁ paratype; A, thoracic shield and tergum 3, lateral view; B, leg 17; C, leg 18; D, E, leg 19 (telopod), caudal and frontal views, respectively. – Scale bar: 0.2 mm (B-E); drawn not to scale (A).

Figure 3. Glomeris colorata sp. n., ♀ paratype from Tunisia, Siliana Gov.; habitus, dorsal view. (Photographed not to scale by I. Muratov).

Figure 4. Glomeris colorata sp. n., ♁ holotype; A, habitus, dorsal view; B, pygidium, caudal view. A schematic, slightly too bright presentation of the colour pattern (del. J.-P. Mauriès).

Figure 6. Glomeris flavomaculata Lucas, 1846, ♁?paralectotype; habitus, dorsal view.(Photographed not to scale by I. Muratov).

Figure 7. Glomeris flavomaculata Lucas, 1846, ♁ paralectotype; A, habitus, dorsal view; B, pygidium, caudal view. A schematic, slightly too bright presentation of the colour pattern (del. J.-P. Mauriès).

Figure 8. Glomeris flavomaculata Lucas, 1846, syntype; habitus, dorsal view. (after Lucas 1849).

Figure 15. Glomeris sublimbata Lucas, 1846, ♁ from Tunisia, Aïn Draham area; habitus, lateral view. (Photographed not to scale by N. Akkari).

Figure 16. Glomeris sublimbata Lucas, 1846, ♁ from Algeria, Guerrounchi Forest (MNHN CC 094); A, habitus, dorsal view; B, pygidium, caudal view. A schematic, slightly too vague presentation of the colour pattern (del. J.-P. Mauriès).

Figure 17. Glomeris sublimbata Lucas, 1846, juvenile ♀ from Algeria, ruisseau des Singes (MNHN CC 094); A, habitus, dorsal view; B, pygidium, caudal view. A schematic, slightly too vague presentation of the colour pattern (del. J.-P. Mauriès).

Figure 20. Glomeris carthaginiensis Schubart, 1953, ♁ topotype; A, B, dorsal and lateral views, respectively, of a rolled animal. (Photographed not to scale by N. Akkari).

Figure 22. Glomeris carthaginiensis Schubart, 1953, ♁ topotype; A, habitus, dorsal view; B, pygidium, caudal view. A schematic, slightly too bright presentation of the colour pattern (del. J.-P. Mauriès).

Figure 24. Glomeris mohamedanica Attems, 1900, ♁ from Tunisia, Nefza; A, habitus, dorsal view; B, pygidium, caudal view. A schematic, slightly too bright presentation of the colour pattern (del. J.-P. Mauriès).

Figure 26. Glomeris punica Attems, 1900, ♁ from Tunisia, Amdoun; A, B, habitus, dorsal and lateral views, respectively. (Photographed not to scale by N. Akkari).

Figure 27. Glomeris punica Attems, 1900, ♁ from Tunisia, Amdoun; A, habitus, dorsal view; B, pygidium, caudal view. A schematic, slightly too vague presentation of the colour pattern (del. J.-P. Mauriès).