Heraeus, Stal, 1862

Pablo M. Dellapé, María C. Melo & Thomas J. Henry, 2016, A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species, Zoological Journal of the Linnean Society 177 (177), pp. 29-134 : 32-37

publication ID	https://doi.org/ 10.1111/zoj.12362
DOI	https://doi.org/10.5281/zenodo.5611101
persistent identifier	https://treatment.plazi.org/id/6F64DB1B-5E43-FFA7-FC7C-F914FDD0FE80
treatment provided by	Plazi
scientific name	Heraeus
status

Treatment

GENUS HERAEUS STÅL, 1862 View in CoL

Type species: Lygaeus (Plociomerus) triguttatus Guérin-Méneville, 1857, by monotypy.

Heraeus Stål, 1862: 314 –315; Stål, 1874: 147; Distant, 1882: 204; Provancher, 1886: 80; Berg, 1892: 162; Lethierry & Severin, 1894: 191; Van Duzee, 1917: 178–179; Barber, 1918: 75; Blatchley, 1926: 389; Barber, 1928: 175; Barber, 1939: 349, 354; Froeschner, 1944: 640; Torre Bueno, 1946: 64, 69–70; Slater & Hurlbutt, 1957: 73; Scudder, 1957: 155; Sweet & Slater, 1961: 339; Slater, 1964: 1081–1082; Slater, 1972: 149; Slater, 1974: 167, 170; Sweet, 1967: 225; Brailovsky, 1979: 549; Harrington, 1980: 108; Brailovsky, 1981: 219; Froeschner, 1981: 43; Froeschner, 1985: 22; Harrington, 1987: 81; Ashlock & Slater, 1988: 227; Slater & Baranowski, 1990 : 130; Baranowski & Slater, 1998: 79; Froeschner, 1999: 244; Baranowski & Slater, 2005: 136; Dellapé 2014: 428, 431; Henry et al. 2015: 488, 499.

Diagnosis

The species included in Heraeus are recognized by the anteriorly extended collar on the ventral region ( Fig. 4 View Figure 4 A), which is a synapomorphy for the genus. All species have: an elongate head with a postocular region longer than the interocular space; the dorsally narrowed and ventrally broadened collar, extending forwards beneath the head ( Figs 1 View Figure 1 , 4 View Figure 4 A); and the aedeagus with a bilobed vesica and a simple conjunctiva, showing relatively small spines in most of the species.

Redescription

Head elongate, postocular distance longer than interocular distance. Buccular juncture V-shaped. Juga rounded. Collar narrow dorsally and broadening ventrally, extending forwards beneath the head. Procoxa with a spine. Profemur with two rows of spines ventrally. Mesofemur with or without spines. Protibia without spines or with small denticles or spines. Tibiae with spiniform setae. Corial margin smooth. Clavus with more than three rows of punctures. Mesepimeron emergent. Evaporative area extensive, not restricted to a small portion of metapleuron surrounding the auricle. Male genitalia: aedeagus without spines or with relatively small spines, with two lobes on vesica lateral to the ejaculatory reservoir.

Distribution

The genus is widely distributed from Canada to Argentina, including the West Indies and the Galapagos Islands. Most of its 39 species are distributed on continental areas. Heraeus plebejus , H. guttatus , and H. triguttatus are also found on at least one island of the West Indies, and some species are known only from the West Indies. Heraeus dominicanus sp. nov. is only known from the Dominican Republic, Heraeus steineri sp. nov. is known from Turks and Caicos Islands, and the islands of the Bahamas, H. caliginosus is known from the Dominican Republic and Haiti, H. hollyae is known from the Dominican Republic, Haiti, and Antigua Island; H. pulchellus is recorded from the Bahamas, Cuba, and the Cayman Islands , and H. concolor is recorded from the Dominican Republic, Haiti, Jamaica, Antigua, Saba, and the Nevis islands. Heraeus pacificus is known only from the Galápagos islands.