Phyllomydas

J. Wilcox & N. Papavero, 1971, The American Genera of Mydidae (Diptera), with the Description of three new Genera and two new Species, Arquivos de Zoologia 21 (2), pp. 41-119 : 104-106

Genus Phyllomydas

Bigot

Phyllomydas Bigot, 1880 : xlvi. Type-species, phylloceT'Us Bigot (mon.).

Head in anterior view one and six-tenths times as broad as high; face at antennae one-half width of head; oral margin situated opposite lower eye margin; gibbosity broad, one and one-half times as broad as high. Probocis and palpi similar to Mydas . Antennal segment 1 cylindrical and nearly as broad as long; 2 about one-third as long as 1 and about three times as broad as long; 3 slender, varying from two and one-half to four and one-half times as long as 1-2; club of some males broadest near base and tapering slightly to apex and two and one-half times as long as 3, fusiform in other males and subequal in length to 3, which is also the usual form of the female club (Figs. 34-37).

Mesonotum with short hairs on lateral margins and dorsocentral stripes. Scutellum bare, usually a few hairs on arms. Postscutellum rugose laterally, bare. Pleura pilose on episternum-L, pteropleura, and metasternum.

Abdominal tergite-la slightly rounded, bare; Lp with numerous short, erect hairs, especially laterally; rather dense, short, recumbent on remaining tergites; bullae (Fig. 54) about three times as long as broad. Sternites with sparse and recumbent hairs, more numerous on 6-8; 0' 8 about one-third lenght of 7. Females with sparse and reci mbent hairs on segments 2-7, numerous erect to slightly retrorse hairs on 8 and a few on 9; tergite 9 fluted and one and one-half times as wide at the apex as at the base; apex with short hairs (Fig. 61). Some species with fine punctures on segments 2-8 and female segment 8 shallowly rugose. Male genitalia (Figs. 122-124): hypandrium completely coalesced with basistyli, resulting structure subquadrate; apical processes of gonopods subrectangular, their apical margin straight; superomedian part of the fused gonopods projected ventrally; aedeagus a conical tube, with lateral apodemes dorsally bent, pointed; epandrial halves and cerci coalescent for a short distance at base; epandrial halves wide, bent in their apical margin, bent part fused to epandrial wall.

Fore and middle legs short and slender. Hind femora moderately swollen, about six times as long as broad; venter with about 14 short tuberculate spines; hairs short. Hind tibiae with a slight ventral keel; apex with a short spur, about one-half width of hind metatarsus, with a long bristle at base and at times a second shorter one. Hind metatarsus about two and one-half times as long as wide and subequal in length to segments 2-3.

Phyllomydas brusii Johnson, male genitalia: 122, ventral view; 123, dorsal view; 124, lateral view.

Alulae with dense fringe of squamose hairs. Wings about- three times as long as broad; second submarginal cell usually closed and petiolate, occasionally narrowly open; posterior crossvein present; ambient vein complete; axillary lobe about as broad as long.

Length: 12-21 mm.

Geographic range: U. S.A. (Arizona and Colorado to Florida), Mexico (Sonora east to Coahuila and south to Jalisco).

Mydas e'ltpolis Seguy: (1928: 142) belongs in Phyllomydas Bigot and this discovery extends the range of the genus much farther south in Mexico than reported previously (Papavero & Wilcox, 1968: 8). The fore and middle tibiae of eupolis are as wide as their femora, the spur on the hind tibiae is slightly curved and one and one-half times the width of hind metatarsus.