Acellomyia paulseni ( Philippi, 1865 )

González, Christian R., 2017, The genus Acellomyia González, a new taxonomic arrangement of its species and descriptions of a new genus and two new species from southern South America (Diptera: Tabanidae: Diachlorini), Zootaxa 4337 (4), pp. 523-539 : 530-532

Acellomyia paulseni ( Philippi, 1865)

( Figs. 4a–h )

Tabanus paulseni Philippi, 1865: 721 .

Archiplatius trianguliferus Enderlein, 1925: 374 . Type-locality: Santiago. HT: ♀ (Berlin). Archiplatius trianguliferus var. ochraceus Enderlein, 1925: 374 . Type-locality: Longaví. HT: ♀ (Berlin). Tabanus (Agelanius) tephrodes Philippi of Kröber, 1930: 151 (misidentify). Dasybasis paulseni Philippi of Stuardo, 1946: 74

Dasybasis (Agelanius) paulseni ( Philippi, 1865) of Coscarón & Philip 1967: 215. Acellomyia paulseni ( Philippi, 1865) of González, 1999: 160.

Type-locality. Unknown. Holotype: ♀ (MNHN).

Diagnosis. Medium-sized species (14.0–15.0 mm). Body color brownish ( Fig. 4a ). Eyes dark brown, with short, sparse ommatrichia. Frons moderately broad and slightly divergent. Basal callus subrectangular. Subcallus with gray-pink setae laterally. Antenna brownish. Maxillary palpus with yellowish pruiniscence, and black and gray setae. Mesonotum dark brown, gray-pink laterally, with short gray and black setae and two gray longitudinal stripes extending to transverse suture. Wing with clouds on the crossveins, without appendix on R4. Abdomen dark brown, bearing short, black setae dorsally; tergites 1–6 reddish brown, with large, pale gray median triangles.

Female. Head. Eyes dark brown, without bands, with short, sparse grayish ommatrichia. Frons with yellowish gray pruiniscence, with short black setae. Frons moderately broad, slightly divergent at the base, frontal index 1.7– 2.2. Basal callus subrectangular, pale brown, touching subcallus and not touching eyes. Ocellar triangle and ocelli absent ( Fig. 4b ). Posterior border of head gray; vertex grayish, with short black and gray setae. Subcallus covered with grayish pruiniscence, and gray-pink setae laterally. Clypeus and genae with grayish pruiniscence, genae with short, gray setae, longer near frontoclypeal suture, clypeus with short, gray setae only laterally. Beard with long gray setae ( Fig. 4c ). Scape of antenna pale brown, with short black setae, pedicel concolorous, with weakly developed dorsal projection and short black setae. First flagellomere dark brown, strongly excavated ( Fig. 4d ), apical flagellomeres concolorous. Maxillary palpus elongated with yellowish pruiniscence and abundant black and gray setae, wider at midlength ( Fig. 4e ). Proboscis dark brown, labella large and fleshy. Thorax. Mesonotum dark brown, gray-pink laterally, with short gray and black setae and two gray longitudinal stripes extending to transverse suture. postpronotal lobes gray-pink, bearing long black setae. Scutellum dark brown on disc, gray-pink, with long gray setae laterally and posteriorly ( Fig. 4f ). Pleura coated with gray-brown pruiniscence, bearing long grayish setae; anepistemum with short black and gray setae. Legs reddish brown, coxae and femora with long gray setae, tibiae with short gray setae, remaining leg segments with short black setae. Wing with clouds on crossveins, Sc and R1 with short setae, without appendix on R4 ( Fig. 4g ). Basicosta bare. Halter dark brown. Squamae concolorous with base of wing. Abdomen. Dark brown, bearing short, black setae dorsally; tergites 1–6 reddish brown, with large, pale gray median triangles, bearing long whitish setae laterally ( Fig. 4h ). Sternites brown, with abundant black setae longer than those on tergites. Terminalia (follows Coscarón & Philip, 1967). Cerci subsemicircular. Tergite 10 with abundant setae. Gonapophysis with distal border slightly projecting and sinuous. Sternite 8 with wide, straight base. Genital fork with wide, slightly concave base. Spermathecal ducts short, spermatheca weakly capitate, with abundant small pores, similar to A. fontanensis .

Male. Length 12.5 mm (n = 2). Similar to female except as follows: eyes with abundant yellowish ommatrichia, upper facets same color as lower facets. Basal flagellomere less excavated. Maxillary palpus cylindrical. Cerci subcircular to triangular. Gonostyli close to A. lanei but bristles shorter. The setulae on the dorsal area of gonocoxite characteristic of this species, do not occur in other species.

Material examined. CHILE: Prov. Valparaíso, 1 ♀, Limache, no other data (paratype) ( MLP); Prov. Cordillera , 2 ♀, RN Río Clarillo, 8.1.1992, C.R. González ( IEUMCE); Prov. Talca, Altos de Vilches , 1280 masl , 3 ♀, 22–29.II.1971 and 1 ♀, 27.Xll.1987, J. Solervicens (IEUMCE); Prov. Linares, 5 ♀, Villalobos, 11.1976, without (IEUMCE); 2 ♀, Parral, II.1976, without coll. (IEUMCE); Prov. Nuble, 1 ♀, Las Trancas, 10–15.XII.1951, L.E. Peña (IEUMCE); 2 ♀, Termas de Chillán, 6.11.1964, Etcheverry & Herrera ( IEUMCE); Prov. Biobío, Laguna Laja 1 ♂, 20.1.1948, Kuschel? ( MLP) ; 1 ♀, 7.I.1968, R. Orca and 1 ♀, 20.1.1948, coll. Illegible ( MLP); Prov. Arauco, Pichinahuel , 3 ♀, Xll.1958–1.1959 and 2 ♀, 31.1.1954, L.E. Peña (MNHN); Nahuelbuta, 4 ♀, 19.1.1985, Ibarra and 1 ♂, II.1988, H. Toro (IEUMCE); Peillem-Pilli Cord., Nahuelbuta, 4 ♀, 13.1.1954, L.E. Peña (MNHN); Prov. Malleco, Pino Hachado, 1800–2000 masl, 4 ♀, 1.1959, L.E. Peña ( MNHN); Liucura , 1800 masl , 7 ♀, 1.1959, L.E. Peña ( MLP); Marimenuco , 1 ♀, 1.1959 and 1 ♀, 1– 18.1.1964, L.E. Peña (MLP); 2 ♀, Cord. Las Raíces, 25– 28.Xll.1967, L.E. Peña (CNC); 1 ♂, Lago Galletué, Cord. Lonquimay, 1.11.1962, L.E. Peña (MNHN); 1 ♀, Termas Tolhuaca, 1.1959, L.E. Peña (MLP); 1 ♀, Altos de Nahuelbuta , 8.I.1930, E.P. Reed ( IEUMCE) ; 1 ♀, Río Blanco , 5–27.111.1951, L.E. Peña ( MLP) ; 2 ♀, W of Angol, 1200 masl, 3.I.1951, Ross & Michelbacher (MLP); l ♀, Angol, 7.I.1930, E.P. Reed (IEUMCE); 7 ♀, 30 km W Angol , 13.11.1967, E.I. Schlinger ( CNC); Prov. Cautín , 1 ♀, Cherquenco, 1.1954, L.E. Peña (MLP); 1 ♀, El Llaima, 26.11.1967, P. Arretz (IEUMCE); Prov. Valdivia, 1 ♀, Pto. Fuy, 5.11.1991, J Solervicens ( IEUMCE); Prov. Osorno, Pto. Puyehue, l ♂, 13–19.IIl.1955 and 7 ♀, 19.1.1969, L.E. Peña ( MLP); l ♀, Puyehue, 4.I.1968, Cekalovic ( MLP); Antillanca , 15 ♀, 12.1.1989 and 9 ♀, 2.11.1992, C.R. González (IEUMCE); 12 ♀, Antillanca, 11– 18.1.1989, S. Coscarón (MLP); 2 ♀, Entre Lagos, Camping Los Copihues, 14.1.1989, S. Coscarón (MLP); Prov. Llanquihue, 2 ♀, Pta. Huano, P.N. Vicente Pérez Rosales, 26.1.1973, J Solervicens (IEUMCE); 1 ♀, C° Derrumbe, P.N. Vicente Pérez Rosales, 20.1.1972, D. Lanfranco (IEUMCE); Prov. Palena, 2 ♀, Palena, I. 1887 without coll. ( MLP); Prov. Aisén , 4 ♀, Pto. Aisén, 22.1.1968, C.W. O’Brien (CNC); Prov. Coihaique, 12 km E. Coihaique, 1 ♀, 22.1.1968 and 1 ♀, 20.1.1968, L. & C.W. O'Brien (CNC); 1 ♀, Lago Escondido, 26 km E. Coihaique, 22.1.1968, L. & C.W. O'Brien (CNC); 5 ♀, Coihaique, 26.1.1977, C. Vivar (MLP); 5 ♀, Balmaceda, 6.1.1961, L.E. Peña ( MLP); No Province data , 4 ♀, Ed C. Reed (MLP), without date; 1 ♀, date, locality and coll. Illegible (MLP); 1 ♀, Sammlung Wolfhülgel; l ♀, V. Dickson, 13.1.1955, Cekalovic (MLP); 4 ♀, no other data ( MLP). ARGENTINA: Prov. Neuquén , 4 ♀, Lago Huechulafquen, 10.1.1968, S. Coscarón (MLP); 4 ♀, Villa Angostura, 18.1.1968, S. Coscarón (MLP); 20 ♀, C° La Atravesada, 50 km W. Zapala, 26.1.1975, L. Pereira (MLP); 2 ♀, Piedra del Águila, 9.Xll.1962, without coll. (MLP); 6 ♀, Lago Lacar, 650 masl, 19.1.1982, M. & P. Gentile (MLP); 8 ♀, Rio Caleufu , 850 masl, 7.1.1981, M. Gentile ( MLP) ; 4 ♀, Cordón Chapelco, Lago Kika, 1750 masl, 24.1.1979, Gentile (MLP); 3 ♀, Moquehue, 1200 masl, 23.I.1976, De Ferraris (MLP); 2 ♀, Quemquemtreu, 710 masl, 19.Xll.1977, Schajovskoi (MLP); 4 ♀, San Martin de Los Andes, Hua-Hum, 24.1.1980, S. Coscarón ( MLP) ; 4 ♀, Lago Queñi, 875 masl, 10.XI.1981, Gentile (MLP); 1 ♀, Quechuquina, SM de Los Andes, 23.1.1978, Coscarón (MLP); 1 ♀, Lago Gutiérrez, Bariloche, without date, J. N. de la Barrera (MLP); 1 ♀, Zapala, 8. Xll. 1920, without coll. ( MLP); Prov. Río Negro , 3 ♀, Valle de las Moscas, 4.I.1975, Coscarón (MLP); 2 ♀, Cerro Catedral, 10.11.1977, Coscarón ( MLP); Prov. Chubut , 2 ♀, Lago Vintter, 18.Il.1961, Coscarón (MLP); 2 ♀, Lago Fontana, 4.11.1961, Coscarón (MLP); 1 ♀, El Triana, 11.11.1961, Coscarón ( MLP); Prov. Santa Cruz , 2 ♀, Calafate, 30.1.1983, Coscarón (MLP); without province: 2 ♀, Paso Carirriñe, 1200 masl, 30.XII.1974, Gentile (MLP).

Distribution. This species occurs in Chile from Quillota to Aysén Province, and in Argentina from Neuquén to Santa Cruz Province.

Comments: the differences that justify its new status and its treatment as a valid species are shown in Table 2.

Figures

FIGURE 4. Female of Acellomyia paulseni (Philippi) a. Habitus (scale bar = 5.0 mm); b. Head in frontal view; c. Head in lateral view; d. Antennae (scale bar = 0.5 mm); e. Maxillary palpus (scale bar = 0.5 mm); f. Mesonotum in dorsal view; g. Wing; h. Abdomen in dorsal view (scale bar = 1 mm).