Graphipterus Latreille, 1802

Renan, Ittai, Assmann, Thorsten & Freidberg, Amnon, 2018, Taxonomic revision of the Graphipterusserrator (Forskal) group (Coleoptera, Carabidae): an increase from five to 15 valid species, ZooKeys 753, pp. 23-82 : 29-30

Graphipterus Latreille, 1802

Stagonopterus Chaudoir, 1871 (type species: Carabus serrator Forskål, 1775)

Graphopterus Agassiz, 1847: 167

Type species.

Carabus variegatus Fabricius, 1792 (= Carabus serrator Forskål, 1775).

Diagnosis.

The Graphipterus serrator group is included in the genus Graphipterus based on the following combination of characters:

Clypeus concave at anterior margin, posteriorly well separated from front; labrum wide and short, with well-developed microsculpture and six setiferous pores. Mandibles broad at the base, sharp and strongly curved at tip; labial and maxillary palps long and slender, glabrous with exception of distal end of segments which bear a few hairs; last palpal segments slightly thicker than penultimate ones.

Pronotum transverse and cordiform, slightly convex, usually ornamented by colored scales at the lateral bead, disc with or without scales. Anterior and posterior angles obtuse.

Scutellum triangular, small and short, often hidden by the pronotal base. Flightless. Elytra wide and oval, slightly convex, coalesced along suture, humeri completely rounded; surface covered by dense or sparse scales, white scales creating longitudinal stripes on the radial field and spots on the disc; apex almost truncate. Pygidium not covered by elytra, last visible tergite with colored scales.

Legs long, usually black or brown, protibia with clypsetae (antenna cleaner) and dark parallel spurs, as long as ¾ of protarsomere 1. Mesotibia with two long and thin not serrated spurs, metatibia with one long and thin not serrated spur and one shorter, wide and obtuse spur. Claws of all legs long and smooth on median margin.