Anypodetus fascipennis, Engel, 1924

Londt, J. G. H., 2000, Afrotropical Asilidae (Diptera) 32. A Revision Of Anypodetus Hermann, 1907 With The Description Of Three New Species (Laphriinae), Annals of the Natal Museum 41, pp. 123-128 : 127-137

2a Wings banded with yellow -brown stained areas, membrane extensively covered with microtrichia; scutellar margin lacking setae; setae of prothoracic coxae and macrosetae of terga black; major mystacal macrosetae largely confined to epistomal margin. Illustrations - Head ( Fig. 1 ), Antenna ( Fig. 4 ), Wing ( Fig. 7 ), Hypopygium ( Figs 16-18 ), Ovipositor ( Fig. 34 ), T2 ( Fig. 39 ), Claws ( Fig. 45 ), Distribution ( Fig. 48 ) .....................................................................................

fascipennis Engel

Synonymy: Anypodetus fascipennis Engel, 1924:103 ;

Anypodetus semirufus Engel, 1924: 104 ; Anypodetus maculipennis Ricardo, 1925: 242 .

Material examined: BOTSWANA: 3♂, Farmer’s Brigade, Serowe , 22 ° 25'S: 26 ° 44'E, 1000 m, 28.xi.1980, P Forchhammer, Malaise trap; 4♂ 15♀, Farmer’s Brigade, Serowe , 2226BD, x.1985 (1♂2♀) - xii.1985 (1♀) - 8.xi.1988 (1♀) - 9.xi.1988 (1♂1♀) - x.1989 (5♀) - xi.1989 (2♂5♀), Forchhammer, Malaise trap; 1♂, Farmer’s Brigade, 6 km SE Serowe , 1300 m, 26.x.1984, Forchhammer, Forestry nursery, A. tortillis woodland; 3♂ 1♀, Serowe , 26.xi.1982 (♂&♀) 21.xii.1982 (♂) 23.x.1982 (♂), Forchhammer, day; 1♀, Palapye [22 ° 33'S: 27 ° 08'E], B.P., 18.x.1923, R Stevenson ( SAMC); NAMIBIA: 1♀, SWA [= South West Africa = Namibia], iii.1923, Mus. Exp.; 1♀, Epupa Falls [16 ° 55'S: 13 ° 10'E], Kunene, 20-22.ii.1994, F Koch ( ZMHB); 1♀, Epupa Falls 17 ° 00'S:13 ° 15'E [use 16 ° 55'S:13:10'E], Kunene, 19-21.ii.1994, H Schumann ( ZMHB); 3♀, Ruacana Falls [17 ° 23'S: 14 ° 15'E], Kunene, 23-24.ii.1994, F Koch ( ZMHB) ; 1♀, Kaross [19 ° 30'S: 14 ° 20'E], ii.1925, Mus. Exped. ( SAMC); 1♂, 27 mi S Tsumeb [19 ° 14'S: 17 ° 43'E], 10.xii.1956; 1♀, Outjo [20 ° 07'S: 16 ° 09'E], i.1925, Mus. Exped. ( SAMC); 1♂ 1♀, 80 mi S Gobabis [22 ° 27'S: 18 ° 58'E], 30.xii.1960, WD Haacke; 1♀, Portsmut, Hakos Mts. , 2416AB, 7.ii.1969, B Lamoral; 1♂, Warmbad [28 ° 27'S: 18 ° 44'E], Koakoveld, ii.1925, Mus. Exped. ( SAMC); SOUTH AFRICA: 1♂ 1♀, Beacon Ranch, 20 km NW Gravelotte [23 ° 57'S: 30 ° 37'E], 17.xi.1978, Brothers & J -Guillarmod; 2♂, Moorddrift [24 ° 17'S: 28 ° 57'E], 7-19.x.1907, CJ Swierstra; 1♂ 1♀, Pretoria [District 25 ° 30'S: 28 ° 10'E], i.1919, Dr Brauns; 1♂, Kaapmuiden [25 ° 32'S: 31 ° 19'E], 30.x.1918, RWE Tucker ( SAMC) ; 1♀, Vryburg [26 ° 57'S: 24 ° 44'E], Griqualand West, x.1939, Mus. Staff ( SAMC) ; 1♂ 1♀, De la Rey [= Delareyville 26 ° 42'S: 25 ° 28'E], i.1917 (♂) - 15.i.1917 (♀), Dr Brauns; 11♂4♀, De la Rey , i.1919, Dr Brauns; 1♂ 1♀, 35 km WNW Upington , 2820BD, 20.iii.1980, J Londt & L Schoeman, roadside vegetation; 1♂ 1♀, 10 miles W Hopetown C.P. [29 ° 37'S: 24 ° 05'E], 27.i.1930, HK Munro [ SANC]; 1♀, Bloemfontein [District 29 ° 10'S: 26 ° 00'E], 12.xii.1920, HK Munro; 1♂, 22 km SE Graaff -Reinet on Pearston Rd. , 32 ° 27'S: 24 ° 38'E, 750 m, 7.xii.1989, J&H Londt, Open Karoo scrub/flowers; 1♀, Merweville [32 ° 40'S: 21 ° 31'E] Dist., ii.1941, H. Zinn ( SAMC); 4♂ 6♀, Willowmore [33 ° 17'S: 23 ° 30'E], ii.1907 (♀) - 20.i.1908 (♀) - 1.i.1909 (♂) - 10.ii.1909 (♀) - xii.1912 (♂) - 1.i.1913 (2♀) - xii.1920 (♀) - 5.i.1922 (♂) - 10.xii.1922 (♂), Dr Brauns; 1♂, Willowmore , 2.i.1913, Dr Brauns ( SAMC); ZIMBABWE: 1♂ 2♀, Hillside [17 ° 50'S: 31 ° 05'E], xi.1922 (♂) - 5.xi.1922 (♀) - 4.ii.1923 (♀), Swinburne & Stevenson; 1♂, Sawmills [19 ° 35'S: 28 ° 02'E], 14.xi.1924, RHR Stevenson; 1♂ 1♀, Bulawayo [20 ° 09'S: 28 ° 35'E], xi.1922 (♂) - 1924 (♀), RHR Stevenson; 2♂ 1♀, Bulawayo , 12.x. & 4.xi.1923, R. Stevenson ( SAMC).

Notes: Type material not seen: A. fascipennis - 1♂, South Africa, Willowmore, 1.1.1907, Dr Brauns & 2♂1♀, Delarey, Dr Brauns, types in ZSMC; A. semirufus - 3♂1♀, South Africa, Willowmore, 1.I. - 5.I.1922, Dr Brauns. Engel (1924) did not indicate the type status of each specimen. If these are syntypes it will be necessary to designate lectotypes for both fascipennis and semirufus . I have seen many specimens from both Willowmore and Deleray collected by Brauns. A. maculipennis - 1♂1♀ syntypes ( maculipennis ), Zimbabwe, Saw Mills (and other material from same locality) apparently in the National Museum of Zimbabwe, Bulawayo. I have seen other material from this locality.

A. fascipennis : Examination of the full series of specimens at my disposal supports acceptance of the synonymy of semirufus Engel and maculipennis Ricardo as suggested by Oldroyd (1974). Although I have not seen Ricardo’s syntypes of maculipennis , I here designate the male as lectotype and the female as paralectotype.

A. fascipennis : The most widely distributed species, known from six climatic regions. Not reported from the higher rainfall areas of the eastern parts of the subcontinent.

Figures

Figs 1-6. Anypodetus species. 1-3. Head, anterior aspect. 1. A. fascipennis Engel (Willowmore ♂). 2. A. leucothrix sp. n. (holotype).3. A. nigrifacies Ricardo (Mkuzi ♂). 4-6.Antenna, outer view of left side.4.A.fascipennis Engel(Willowmoreơ). 5.A.nigrifacies Ricardo(Mkuziơ). 6.A.macroceros sp. n. (holotype). Scale lines = 1 mm.

Figs 7-9. Anypodetus species, wing venation. 7. A. fascipennsis Engel (Willowmore ♀). 8. A. nigrifacies Ricardo (Mkuzi ♂). 9. A. leucothrix sp. n.(holotype). Scale lines = 1 mm.

Figs 10-21. Anypodetus species, hypopygia. 10-12. A. arachnoides Oldroyd (6 km N Vivo). 13-15. A. fasciatus Hermann (Serowe). 16-18. A. fascipennis Engel (Willowmore). 19-21. A. leucothrix sp. n. (paratype). Scale lines = 1 mm.

Figs 34-36. Anypodetus species, ovipositor, ventral aspect. 34. A. fascipennis Engel (Willowmore). 35. A. nigrifacies Ricardo (Mkuzi). 36. A. unicolor Oldroyd (20 mi S Ndumu). Scale line = 1 mm.

Figs 37-44.Anypodetus species,secondabdominaltergum,distributionofpruinescence(unhatched).37.A. arachnoides Oldroyd.38.A.fasciatus Hermann.39.A.fascipennis Engel.40.A.leucothrix sp.n. 41.A.macroceros sp.n.42.A.nigrifacies Ricardo.43.A.phalaros sp.n.44.A.unicolor Oldroyd. Scale line = 1 mm.

Figs 45-47. Anypodetus species. 45. A. fascipennis Engel, tip of left hind tarsus. 46. A. unicolor Oldroyd, egg dissected from female abdomen. 47. A. leucothrix sp. n., scutellum, dorsal view. Scale lines = 1 mm.

Figs 48-49. Anypodetus species distribution. 48. A. fascipennis Hermann (closed circles), A. leucothrix sp. n. (closed triangles), A. nigrifacies Ricardo (closed squares), A. phalaros sp. n. (open circles). 49.A.arachnoides Oldroyd(closedtriangles),A.fasciatus Hermann(closedcircles),A.macroceros sp. n. (open circle), A. unicolor Oldroyd (closed squares).